At the time of sporangial formation the protoplasm with numerous nuclei streams into the swollen end of the sporangiophore and there becomes cut off by a cell-wall to form the sporangium.
When the threads reach the air they branch in a tree-like manner, and each branch (sporangiophore) carries one or more ovate sporangia, as shown at E, E, E, which fall off and are carried by the wind.
(All except d represent vertical sections of sporangiophore or sorus.) canal the spermatozoid, which in the Ferns has been shown to be attracted by reason of its positive irritability to malic acid, passes and fuses with the ovum.
From each segment, near its base, a stalked peltate sporangiophore arose; this bore four sporangia, which hung parallel to the stalk.
Other views of the nature of the Psilotaceous synangium are, however, possible, and indeed the existence of both simple and complicated sporangiophores in the Sphenophyllaceae leaves the question open as to whether the synangium in existing Psilotaceae is a relatively simple type of sporangiophore which has persisted unaltered or is the result of reduction from a more elaborate structure.
This singular arrangement has suggested doubts as to the correctness of the current interpretation of the Equisetaceous sporangiophore as a modified leaf Sm ' 'CI= B A (After Renault.
Each sporangiophore is traversed throughout its length by a vascular bundle connected with that which supplies the subtending bract.
Dawsoni in the fact that each sporangiophore bears two sporangia, attached to a distal expansion approaching the peltate scale of the Equisetales.
It is thus proved that the sporangiophore is not a mere sporangial stalk, but a distinct organ, in all probability representing a ventral lobe of the subtending bract.
There is good reason to believe that the ventral synangium of the Psiloteae corresponds to the ventral sporangiophore with its sporangia in the Sphenophyllales.