or many protoxylems. When the protoxylem strands are situated at the periphery of the stele, abutting on the pericycle, as in all roots, and many of the more primitive Pteridophyte stems, the stele is said to be exarch.
When the protoxylems have an intermediate position the stele is inesarch (many Pteridophytes and some of the more primitive Phanerogams).
The protoxylems and the phloem strands are developed alternately, just within the outer limit of the young cylinder.
The anatomy of Lycopodium presents considerable variety in detail, but the stem is always monostelic and the development of the xylem centripetal, the protoxylems being situated at the periphery of the stele; pericycle and endodermis surround the stele, and the wide cortex may be more or less sclerenchymatous.
While the stele of the primary root in both Gymnosperms and Angiosperms is usually diarch or tetrarch, the large primary root-steles of many adventitious roots are frequently polyarch, sometimes with a very large number of protoxylems. Such a stale seldom has the centre filled up with xylem, this being replaced by a large-celled pith, so that a siphonostelic structure is acquired (fig.
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