Their structure has undergone little degeneration in connexion with this habit, and may be compared organ for organ with that of the Planarians.
Supposing the tube to be completely introverted and to commence its eversion, we then find that eversion may take place, either by a forward movement of the side of the tube near its attached base, as in the proboscis of the Nemertine worms, the pharynx of Chaetopods and the eye-tentacle of Gastropods, or by a forward movement of the inverted apex of the tube, as in the proboscis of the Rhabdocoel Planarians, and in that of Gastropods here under consideration.
The interesting little Rhodope veranyii, which has no odontophore, has been associated by systematists both with these simplified Opisthobranchs and with Rhabdocoel Planarians.
Temnocephaloidea (see appendix tO Planarians).
The concentration of nervous matter and ganglionic substance at the oral end of Trematodes is equivalent to the " brain " of the Planarians, but the similar thickening in the scolex of Cestodes is by no means so certainly to be called by that name.
The last structural peculiarity of the group is the absence of the functions of regulation and reparation which are so highly developed in the more primitive Planarians.
From these are given off at irregular intervals short lateral branches, each of which terminates in a flame-cell (f) precisely similar in structure to the flame-cells found in Planarians, Trematodes and Cestodes; here as there the question whether they are open to the body cavity or not must probably be answered in the negative.
PLATYELMIA, a phylum of the animal kingdom which comprises three classes, the Planarians, Trematodes and Cestodes.
The Planarians are free-living animals, the Trematodes are parasitic upon and within animals, and the Cestodes are wholly endoparasitic.
Within this muscular tube lies a parenchymatous tissue which may be uniform (Cestodes) or differentiated into a central or digestive, and a peripheral portion (some Turbellaria), or finally the central portion becomes tubular and forms the digestive sac (Trematodes), while the peripheral portion is separated from it by a space lined in some forms by a flattened epithelium (most Planarians).
As the Turbellaria (Planarians) are the most primitive division of the Platyelmia, the problem of the affinities of this phylum resolves itself into that of the relationships of the Turbellaria.
Some Opisthobranchia are striking examples of degeneration (some Nudibranchia), having none of those regions or processes of the body developed which distinguish the archaic Mollusca from such flat-worms as the Dendrocoel Planarians.
Planaria (see PlanariaNS).
The class as a whole is linked to the Turbellaria not only by its similarity of structure, but by the intermediation of the singular class the Temnocephaloidea (see Planarians), which in habit and in organization form an almost ideal annectant group.
The Trematodes are somewhat modified in accordance with their ectoor endoparasitic life, but they exhibit such a close similarity of structure with the Turbellaria that their origin from Planarians can hardly be doubted, and indeed the Temnocephaloidea (see Planarians) form an almost ideal annectant group linking the ectoparasitic Trematodes and Rhabdocoel Planarians.
The so-called flatworms (Platyelmia, q.v.), including the Planarians, Flukes (see Trematodes), Cestodes (see Tapeworm) and the curious Mesozoa, are no doubt related.
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