Immediately above the notochord there lies another FIG.
As already stated, the notochord extends beyond the mouth to the tip of the rostrum.
C. Labyrinthodonta, with simple biconcave vertebral disks, very slightly pierced by a remnant of the notochord and supporting the loosely articulated neural arch.
It has, in fact, become metamorphosed into a resistant supporting structure resembling in some respects the notochord of the true Chordata, but probably not directly comparable with the latter structure, being related to it solely by way of substitution.
- anp, Anterior neural pore; be, rudiment of buccal skeleton; c, cilia; cb, ciliated band; cc, ciliated groove; cm, cilia at margin of mouth; gl, external opening of club-shaped gland; Hn, Hatschek's nephridium; lm, left metapleur; n, notochord; pp, praeoral pit; ps, primary gill-slits, I, 5, and 13; rm, right metapleur showing through.
a, Cavity surrounding fin ray; a', fin ray; b, muscular tissue of myotome; c, nervecord; d, notochord; c, left aorta; f, thickened ridges of epithelium of praeoral chamber (Rader organ); g, coiled tube lying in a coelomic space on right side of praeoral hood, apparently an artery; h, cuticle of notochord; i, connective-tissue sheath of notochord; k, median ridge of skeletal canal of nerve-cord; 1, skeletal canal protecting nerve-cord; m, inter-segmental skeletal septum of myotome; n, subcutaneous skeletal connective tissue; o, ditto of metapleur (this should be relatively thicker than it is); q, subcutaneous connective tissue of ventral surface of atrial wall (not a canal, as supposed by Stieda and others); r, epiblastic epithelium; s, gonad-sac containing ova; t, pharyngeal bar in section, one of the "tongue" bars alternating with the main bars and devoid of pharyngo-pleural fold and coelom; v, atrio-coelomic funnel; w, socalled "dorsal" coelom; x, lymphatic space or canal of metapleur; y, sub-pharyngeal vascular trunk; z, blood-vessel (portal vein) on wall of hepatic caecum; aa, space of atrial or branchial chamber; bb, ventral groove of pharynx (anteriorly this takes the form of a ridge); cc, hyperbranchial groove of pharynx; dd, lumen or space of hepatic caecum; ee, narrow coelomic space surrounding hepatic caecum; $, lining cell-layer of hepatic caecum; gg, inner face of a pharyngeal bar clothed with hypoblast, the outer face covered with epiblast (represented black); hh, a main pharyngeal bar with projecting pharyngeal fold (on which the reference line rests) in section, showing coelomic space beneath the black epiblast; ii, transverse ventral muscle of epipleura; kk, raphe or plane of fusion of two down-grown epipleura; 11, space and nucleated cells on dorsal face of notochord; mm, similar space and cells on its ventral face.
I, 2), in which the spinal cord rests on the notochord, which persists uninterrupted and is surrounded by three bony elements in addition to the neural arch: a so-called pleurocentrum on each side, which appears to represent the centrum proper of reptiles and mammals, and an intercentrum or hypocentrum below, which may extend to the neural arch, and probably answers to the hypapophysis, as it is produced into chevrons in the caudal region.
A ventral vessel occurs on the anterior side of the metasome and forms a loop extending down the entire length of the stalk, while a " heart " projects into the cavity of the pericardium, probably connected on the ventral side of the notochord with the ventral vessel, and on its dorsal side with the dorsal vessel.
The ordinal name for the genera and species of Amphioxus is Cephalochorda, the term referring to the extension of the primary backbone or notochord to the anterior extremity of the body; the family name is Branchiostomidae.
In Asymmetron the caudal region is remarkable for the curious elongation of the notochord, which is produced far beyond the last of the myotomes.
First, its dorsal wall (which is grooved to form the hyperpharyngeal groove) is closely adherent to the sheath of the notochord; and secondly, the pharynx is attached through the intermediation of the primary bars.
- As has already been indicated, a solid subcylindrical elastic rod, the notochord, surrounded by a sheath of laminar connective tissue, the cordal sheath, lies above the alimentary canal in contact with its dorsal wall, and extends beyond it both in front and behind to the obtusely pointed extremities of the body.
This notochord represents the persistent primordial skeletal axis which, in the higher Craniata (though not so in the lower), gives way by substitution to the segmented vertebral column.
It is the central nervous system, and contains within itself the elements of the brain and spinal marrow of higher forms. The neurochord tapers towards its posterior end, where it is coextensive with the notochord, but ends abruptly in front, some distance behind the tip of the snout.
- a, Atrium; al, alimentary canal; y blood-vessel; cv, cerebral vesicle; df, dorsal section of myocoel (= fin spaces); e, " eyespot"; end, endostyle; gl, club-shaped gland; lm, edge of left metapleur; m, lower edge of mouth; n, notochord; nt, pigmented nerve tube; ps, primary gill-slits, I, 9, and 14; rc, renal cells on atrial floor; rm, edge of right metapleur; so, sense organ opening into praeoral pit; ss, thickenings, the rudiments of the row of secondary gill-slits.
After invagination is completed, the embryo begins to elongate, the blastopore becomes narrower, and the dorsal wall of the gastrula loses its convexity, and becomes flattened to form the dorsal plate, the outer layer of which is the primordium of the neurochord and the inner layer the primordium of the notochord.
(Adapted from Hatschek.) pc, Praechordal head-cavity of embryo; cc, collar-cavity (first somite); my, mesodermic somites (myocoelomic or archenteric pouches); ch, notochord with the neural tube (neurochord) lying upon it; np, anterior neuropore; ne, position of posterior neurenteric canal.
Embolomeri, with the centra and intercentra equally developed disks, of which there are thus two to each neural arch; these disks perforated in the middle for the passage of the notochord.
Microsauria, nearest the reptiles, with persistent notochord completely surrounded by constricted cylinders on which the neural arch rests.
Branchiosauria, nearest to the true batrachians; with persistent non-constricted notochord, surrounded by barrel-shaped, bony cylinders formed by the neural arch above and a pair of intercentra below, both these elements taking an equal share in the formation of a transverse process on each side for the support of the rib.
In some the notochord remains for a long time exposed along the ventral surface, and, owing to the absence of cartilaginous formation around it, disappears without ever becoming invested otherwise than by a thin elastic membrane; it can be easily stripped off below the vertebrae in larval specimens on the point of metamorphosing.
In others, which represent the perichordal type, the greater share of the formation of the whole vertebra falls to the (paired) dorsal cartilage, but there is in addition a narrow ventral or hypochordal cartilage which fuses with the dorsal or becomes connected with it by calcified tissue; the notochord is thus completely surrounded by a thick sheath in tadpoles with imperfectly developed limbs.
They are the only pelagic forms that have a notochord and they retain it throughout their lives.
The development of the spine starts with a band of fibrous tissue called the notochord.
notochord differentiation, leading to embryos of significantly diminished stature.
2, nch.), growing forwards in the septum between the proboscis-cavity and the collar-cavities, and supported dorsally by the median mesentery of the collar, is the representative of the so-called notochord or stomochord of Balanoglossus; and if the view that this organ is really a notochord is well founded, it may be regarded as the homologue of the anterior end of the Vertebrate notochord.
a, Olfactory ciliated pit on animal's left side, its wall confluent with substance of nerve tube; b, pigment spot (rudimentary eye) on anterior termination of nerve tube; c, first pair of nerves in section; d, fin ray;, e, myotome; f, notochord; g, space round myotome (?artifact or coelom); h, subchordal canal (?
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