Others however (inclusive of Tritylodon and Microlestes, if they be really mammals), seem nearer to the Monotremata; and the question has yet to be decided whether placentals and marsupials on the one hand, and monotremes on the other are not independently derived from reptilian ancestors.
In common with the other monotremes, the male echidna has its heel provided with a sharp hollow spur, connected with a secreting gland, and with muscles capable of pressing the secretion from the gland into the spur.
Monotremes (2 species) and marsupials (4 families and 44 species) predominate, but are not abundant.
Although the present article does not discuss mammalian osteology in general (for which see Vertebrata), it is interesting to notice in this connexion that the primitive condition of the mammalian tympanum apparently consisted merely of a small and incomplete bony ring, with, at most, an imperfect ventral wall to the tympanic cavity, and that a close approximation to this original condition still persists in the monotremes, especially Ornithorhynchus.
Adult monotremes are in like case, although the duckbilled platypus (Ornithorhynchus) has teeth when young on the sides of the jaws.
Very different, on the other hand, is the condition of things in Australia and Papua, where marsupials (and monotremes) are the dominant forms of mammalian life, the placentals being represented (apart from bats, which are mainly of an Asiatic type) only by a number of more or less aberrant rodents belonging to the mouse-tribe, and in Australia by the dingo, or native dog, and in New Guinea by a wild pig.