VII, Usually considered to be the tergum of the genital somite, but suggested by Pocock to be that of the other [According to the system of numbering explained in the text, if VII is the tergum of the praegenital somite (as is probable) it should be labelled Prg without any number, and the somites VIII to XIII should be lettered 1 to 6, indicating that they are the six normal somites of the mesosoma; whilst XV to XVIII should be replaced by the numbers 7 to 12 - an additional suppressed segment (making up the typical six) being reckoned to the metasomatic fusion.] (From Lankester, Q.
Passing on now from the mesosoma we come in Scorpio to the metasoma of six segments, the first of which is broad whilst the rest are cylindrical.
42) the six somites of the mesosoma are not fused to form a carapace at an early stage, and they are followed by three separately marked metasomatic somites; the other three somites of the metasoma have disappeared in Limulus, but are represented (From Lankester, loc. cit.) by the unsegmented prae-anal region.
VIII to XIII, The six somites of the mesosoma, each with a movable pleural spine and a pair of dorsal entopophysis or muscle-attaching ingrowths.
7, excepting that VII is here certainly the tergum of the first somite of the mesosoma - the genital somite - and is not a survival of the embryonic praegenital somite.
Perhaps the most important general agreement of Scorpio compared with Limulus and the Eurypterines is the division of the body into the three regions (or tagmata) - prosoma, mesosoma and metasoma - each consisting of six segments, the prosoma having leg-like appendages, the mesosoma having foliaceous appendages, and the metasoma being destitute of appendages.
The soft integument and limbs of the mesosoma have been removed as well as all the viscera and muscles, so that the inner surface of the terga of these somites with their entopophyses are seen.
The first segment of the mesosoma of Scorpio and Limulus thus remains the first segment, and can be identified as such throughout the Eu-arachnida, carrying as it always does the genital apertures.
We shall, therefore, ignoring the ocular somite, speak of the first, second, third, fourth, fifth and sixth legbearing somites of the prosoma, and indicate the appendages by the Roman numerals, I, II, III, IV, V, VI, and whilst ignoring the praegenital somite we shall speak of the first, second, third, &c., somite of the mesosoma or opisthosoma (united mesosoma and metasoma) and indicate them by the Arabic numerals.
VIIgo, The genital somite or first somite of the mesosoma with the genital operculum (a fused pair of limbs).
In each somite of the mesosoma is a small, free entosternite having a similar position, but below or ventral to the nerve cords, and having a smaller number of muscles attached to it.
In the mesosoma the approximation had occurred before the entosternites were formed.
In Limulus small entosternites are found in each somite of the appendage-bearing mesosoma, and we find in Scorpio, in the only somite of the mesosoma which has a welldeveloped pair of appendages, that of the pectens, a small entosternite with ten pairs of muscles inserted into it.
Intermediate somites forming a mesosoma occur, but tend to fuse superficially with the metasomatic carapace or to become co-ordinated with the somites of the metasoma, whether fused or distinct to form one region, the opisthosoma (abdomen of authors).
In the most highly developed forms the two anterior divisions (tagmata) of the body, prosoma and mesosoma, each exhibit six pairs of limbs, pediform and plate-like respectively, whilst the metasoma consists of six limbless somites and a post-anal spine.
In this genus ten free somites (mesosoma) occur between the prosomatic and metasomatic carapaces.
This telson may enlarge, it may possibly even become internally and sternally developed as partially separate somites, and the tergum may remain without trace of somite formation, or, as appears to be the case in Limulus, the telson gives rise to a few well-marked somites (mesosoma and two others) and then enlarges without further trace of segmentation, whilst the chitinous integument which develops in increasing thickness on the terga as growth advances welds together the unsegmented telson and the somites in front of it, which were previ ously marked by separate tergal thickenings.
(After Beecher.) ancestor of the grade) grouped in three regions of six - (a) the " prosoma " with palpiform appendages, (b) the " mesosoma " with plate-like appendages, and (c) the " metasoma " with suppressed FIG.
A somite placed between the prosoma and mesosoma - the prae-genital somite - appears to have belonged originally to the prosomatic series (which with its ocular prosthomere and palpi FIG.
(After Beecher, from Zittel.) retained as a rudimentary, separate, detached somite in front of the mesosoma, or disappears altogether (excalation).
Concentration of the organ-systems by fusion of neighbouring regions (prosoma, mesosoma, metasoma), pre viously distinct, has frequently occurred, together with obliteration of the muscular and chitinous structures indicative of distinct somites.
- Nomomeristic Arachnids, in which the somites corresponding to mesosoma and metasoma have entirely aborted.
- These start from highly developed and specialized aquatic branchiferous forms, exhibiting a prosoma with six pediform pairs of appendages, an intermediate prae-genital somite, a mesosoma of six somites bearing lamelliform pairs of appendages, and a metasoma of six somites devoid of appendages, and the last provided with a post-anal spine.
Not freeswimming, none of the prosomatic appendages modified to act as paddles; segments of the mesosoma and metasoma (= opisthosoma) not more than ten in number, distinct or coalesced.
- Free-swimming forms, with the appendages of the 6th or 5th and 6th pairs flattened or lengthened to act as oars; segments of mesosoma and metasoma (= opisthosoma), twelve in number.
The appendages of the mesosoma generally suppressed; in the more primitive forms one or two pairs may be retained as organs subservient to reproduction or silkspinning.
- The primitive distinction between the mesosoma and the metasoma retained, the latter consisting of six somites and the former of six somites in the adult, each of which is furnished during growth with a pair of appendages.
- The primitive distinction between the mesosoma and the metasoma wholly or almost wholly obliterated, the two regions uniting to form an opisthosoma, which never consists of more than twelve somites and never bears appendages or breathing-organs behind the 4th somite.
- Appendages of 1st pair bisegmented, without poison gland; of 2nd pair prehensile, their basal segments underlying the proboscis, and furnished with sterno 1 to i 1, Somites of the opisthosoma (mesosoma plus metasoma).
- The possibility of another interpretation of the anterior somites of the mesosoma and the prae-genital somite must be borne in mind.
In higher specialized forms these branchial processes become first of all limited to five segments of the mesosoma, then sunk beneath the surface as pulmonary organs, and finally atrophied, their place being taken by a well-developed tracheal system.