D, Embryo with lateral torsion and an endogastric shell.
The name expresses the most universal character of the class, the importance of which was first noticed by John Ray, namely, the presence of a pair of seed-leaves or cotyledons, in the plantlet or embryo contained in the seed.
The parasite effects a lodgment in the host either by invading it as a free-swimming planula, or, apparently, in other cases, as a spore-embryo which is captured and swallowed as food by the host.
Such cannulated cells are characteristic of the nephridia of many worms, and the organs thus formed in the embryo Limnaeus are embryonic nephridia.
The seeds are minute and innumerable; they contain a small rudimentary embryo surrounded by a thin loose membraneous coat, and are scattered by means of hygroscopic hairs on the inside of the valves which by their movements jerk out the seeds.
If we go back to the first instance cited, the embryo in the seed and its development during germination, we can ascertain what is necessary for its life by inquiring what are the materials which are deposited in the seed, and which become exhausted by consumption as growth and development proceed.
Areas are found on the foot of the embryo Pulmonate Limax and on the yolk-sac (distended foot-surface) of the Cephalopod Loligo.
The rows of cells from which the laticiferous vessels are formed can be distinguished in many cases in the young embryo while still in the dry seed (Scott), but the latex vessels in process of formation are more easily seen when germination has begun.
The embryo consists of an axis bearing two or more cotyledons and ending below in a radicle; it lies in a generally copious food-storing tissue (endosperm) which is the remains of the female prothallus.
The body thus formed ment of is called the embryo, and this develops into the adult Primary plant, not by continued growth of all its parts as in an animal, but by localization of the regions of cell-division and growth, such a localized region being called a growing-point.
It is absent in the Ratitae, which from this feature have received their name, but considerable traces of a cartilaginous keel occur in the embryo of the ostrich, showing undeniably that the absence of a keel in the recent bird is not a primitive, fundamental feature.
Size compared with the thorax or abdomen, but in the embryo it On the whole it seems likely that the endoderm is represented in forms a much larger portion of the body than it does in the adult.
It is absurd to call the larva of a newt or of a Caecilian a tadpole, nor is the free-swimming embryo of a frog as it leaves the egg a tadpole.
B, Embryo with ventral flexure of the intestine.
In cases where the development of the embryo is advanced at the resting period, traces run from the cotyledons and determine the symmetry of the stele of the primitive axis, the upperpart of which often shows stem-structure, in some respects at least, and is called the hypocoty- ledonary stem or hypocotyl, while the lower part is the primary root .~-,
C, Embryo with ventral flexure and exogastric shell.
The preconchylian invagination or shell-gland is formed in the embryo behind the velum, on the surface opposite the blastopore.
The foot is always simple, with its flat crawling surface extending from end to end, but in the embryo Limnaea it shows a bilobed character, which leads on to the condition characteristic of Pteropoda.
- Embryo of Limnaeus stagnalis, at a stage when the Trochosphere is developing foot and shell-gland and becoming a Veliger, seen as a transparent object under slight pressure.
When the middle and hinder regions of the blastopore are closing in, an equatorial ridge of ciliated cells is formed, converting the embryo into a typical trochosphere.
In the case of the great grey kangaroo, for instance, the period of gestation is less than forty days, and the newly-born embryo, which is blind, naked, and unable to use its bud-like limbs, is little more than an inch in length.
If the three principal organ-systems of the medusa, namely mouth, tentacles and umbrella, be considered in the light of phylogeny, it is evident that the manubrium bearing the mouth must be the oldest, as representing a common property of all the Coelentera, even of the gastrula embryo of all Enterozoa.
After fertilization the female cell, now called the oospore, divides and part of it develops into the embryo (new sporophyte), which remains dormant for a time still protected by the ovule which has developed to become the seed.
When the young sporophyte first begins its independent lifewhen, that is, it exists in the form of the embryo in the seedits living substance has no power of utilizing the simple inorganic compounds spoken of.
The Pulmonata have a straight visceral nerve-loop, usually no operculum even in the embryo, and a multidenticulate radula, the teeth being equi-formal; and they are hermaphrodite.
It is not true, for example, that a fish is a reptile arrested in its development, or that a reptile was ever a fish; but it is true that the reptile embryo, at one stage of its development, is an organism which, if it had an independent existence, must be classified among fishes; and all the organs of the reptile pass, in the course of their development, through conditions which are closely analogous to those which are permanent in some fishes.
The female is viviparous, and the young, which, unlike the parent, are provided with a long tail, live free in water; it was formerly believed from the frequency with which the legs and feet were attacked by this parasite that the embryo entered the skin directly from the water, but it has been shown by Fedschenko, and confirmed by Manson, Leiper and others, that the larva bores its way into the body of a Cyclops and there undergoes further development.
If the embryo is set free as a free-swimming, so-called planula-larva, in the blastula, parenchymula, or gastrula stage, then a free actinula stage is not found; if, on the other hand, a free actinula occurs, then there is no free planula stage.
And of the muscid flies, an anterior and a posterior endodermThe embryo thus becomes transferred to the dorsal face of the egg, rudiment both derived from the " endoblast " become apparent but at a later stage it undergoes reversion to its original ventral at an early stage, in close association with the stomodaeum and position.
The embryo consists of an axis bearing one (Monocotyledons) or two (Dicotyledons) cotyledons, which protect the stem bud (plumule) of the future plant, and ending below in a radicle.
In fact, there is a period when, as Aristotle long ago said, the embryo of the highest animal has the form of a mere worm, and, devoid of internal and external organization, is merely an almost structureless lump of polype-substance.
He showed that all the organs of plants are built up of cells, that the plant embryo originates from a single cell, and that the physiological activities of the plant are dependent upon the individual activities of these vital units.
The external form of the embryo goes through the same changes as in other Gastropods, and is not, as was held previously to Lankester's observations, exceptional.
Chun and Woltereck, on the other hand, regard the stem as a stolo prolifer arising from the aboral pole, that is to say, from the ex-umbrella, similar to that which grows out from the ex-umbral surface of the embryo of the Narcomedusae and produces buds, a view which is certainly supported by the embryological evidence to be adduced shortly.
Later, the axis branches by the formation of new growing-points, and in this way the complex system of axes forming the body of the ordinary vascular plant is built up. In the flowering plants the embryo, after developing up to a certain point, stopf growing and rests, enclosed within the seed.
In germination of the seed the root of the embryo (radicle) grows out to get a holdfast for the plant; this is generally followed by the growth of the short stem immediately above the root, the so-called "hypocotyl," which carries up the cotyledons above the ground, where they spread to the light and become the first green leaves of the plant.
It has been ascertained that the nephridia of Oligochaeta are preceded in the embryo by a pair of delicate and sinuous tubes, also found in the Hirudinea and Polychaeta, which are larval excretory organs.
The embryo of a moth, a dragon-fly point.
(After Robert.) A, Embryo without flexure.
- Embryo of Cavolinia tridentate.