ascophorous, producing asci; ascospore, the spore (or sporule) developed in the ascus; ascogonium, the organ producing it, &c.
He showed that the archicarp consisted of two parts, a lower coiled portion, the ascogonium, and an upper portion, the trichogyne, which projected from the thallus.
Only when a spermatium was found attached to the trichogyne did the further development of the ascogonium take place.
In some forms we find definite male and female sexual organs (Sphaerotheca, Pyronema, &c.), in others the antheridium is abortive or absent, but the ascogonium (oogonium) is still present and the female nuclei fuse in pairs (Lachnea stercorea, Humaria granulata, Ascobolus furfuraceus); while in other forms ascogonium and antheridium are both absent and fusion occurs between vegetative nuclei (Humaria rutilans, and probably the majority of other forms).
6, The multicellular ascogonium 3, Passage of the antheridial derived by division from the nucleus towards that of the oogonium; the terminal cell oogonium.
When the ascogonium (female organ) is present the ascogenous hyphae arise from it, with or without its previous fusion with an antheridium.
Concerning the second question, the recent investigations of Buchner ascogenous hyphae with their asci represent the sporophyte since they are derived from the fertilized ascogonium.
The matter is complicated by the apogamous transition from gametophyte to sporophyte in the absence of the ascogonium; also by the fact that there are normally two fusions in the life-history as mentioned earlier.
The latter arise from the crown of a spirally coiled archicarp (bearing an ascogonium at its end) and a straight antheridium.
D, The same, beginning to be as, Ascogonium.
are coenogametes), but that the antheridium disorganizes without passing its contents into the ascogonium.
ascogonium and antheridium contain numerous nuclei; they are to be looked upon as gametangia in which there is no differentiation of gametes, and since they act as single gametes they are termed coenogametes.
The cytological details of development of the perithecia are not well known; most of them appear to develop their ascogenous hyphae in an apogamous way without any connexion with an ascogonium.
In relation to the view that the spermatia are sexual cells, or at least were primitively so, it must be pointed out that although the actual fusion of the spermatial nucleus with a female nucleus has not been observed, yet in a few cases the spermatia have been seen to fuse with a projecting portion (trichogyne) of the ascogonium, as in Collema and Physcia, and there is very strong circumstantial evidence that fertilization takes place (see later in section on development of ascocarp).
ascogonium and antheridium contain a number of nuclei (i.e.
In some forms as in Ascobolus the ascogonium is multicellular, the various cells communicating by pores in the transverse walls (fig.
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