It remains to be seen if the ovuliferous cone in the centre of the flower represents simply a functionless gynoecium, as in Welwitschia and abnormal cones of certain Coniferae, or if the flowers were hermaphrodite, with both male and female organs fully developed.
This fact suggests the possibility that the flowers described by Mr Wieland, in which the male organs are mature and the gynoecium is composed of very short and immature ovuliferous stalks and interseminal scales, are not essentially distinct from those which have lost the staminate leaves FIG.
It is interesting that no monstrous cycadean cone has been described in which ovuliferous and staminate appendages are borne on the same axis: in the Bennettitales (see Palaeobotany: Mesozoic) flowers were produced bearing on the same axis both androecium and gynoecium.
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