The great order of Ungulata is represented by various forms of sheep, as many as ten or twelve wild species of Ovis being met with in the mountain chains of Asia; and more sparingly by several peculiar forms of antelope, such as the saiga (Saiga tatarica), and the Gazella gutturosa, or yellow sheep. Coming to the deer, we also meet with characteristic forms in northern Asia, especially those belonging to the typical genus Cervus.
As ancestors of the Artiodactyle section of the Ungulata, we may look to forms more or less closely related to the North American Lower Eocene genera Mioclaenus and Pantolestes, respectively typifying the families Mioclaenidae and Pantolestidae.
All Ungulata probably originated from Condylarthra.
The Perissodactyla have been brigaded with the Artiodactyla to form the typical group of the ungulates, under the name of Diplarthra, or Ungulata Vera, and the features distinguishing the combined group from the less specialized members of the order Ungulata will be found under the heading of that order.
Orders: Insectivora, Chiroptera, Dermoptera, Edentata (Sub-orders: Xenarthra, Pholidota, Tubulidentata), Rodentia (Sub-orders: Duplicidentata, Simplicidentata), Tillodontia, Carnivora (Sub-orders: Fissipedia, Pinnipedia, Creodonta), Cetacea (Sub orders: Archaeoceti, Odontoceti, Mystacoceti), Sirenia, Ungulata (Sub-orders: Proboscidea, Hyracoidea, Barypoda, Toxodontia, Amblypoda, Litopterna, Ancylopoda, Condylarthra, Perissodactyla, Artiodactyla), Primates (Sub-orders: Prosimiae, Anthropoidea).
UNGULATA, the name of an order of placental mammals in which the terminal joints of the toes are usually encased in solid hoofs or covered with broad hoof-like nails, while the molar (and not unfrequently some or all of the premolar) teeth have broad tuberculated crowns adapted for crushing vegetable substances.
In the typical Ungulata or Diplarthra, the feet are never plantigrade, and the functional toes do not exceed four - the inner digit being suppressed, Right Fore Foot of Indian at all events in all forms which Elephant.
ARSINOITHERIUM (so called from the Egyptian queen Arsinoe), a gigantic horned mammal from the Middle Eocene beds of the Fayum, Egypt, representing a sub-order of Ungulata, called Barypoda.
The Ungulata are represented by the chamois (Rupicapra tragus) and the bouquetin or steinbock (Capra ibex).
We are dependent upon the Carnivora, Rodentia, Ungulata and Marsupialia for our supplies of furs, the first two classes being by far of the greatest importance.
The Carnivora include bears, wolverines, wolves, raccoons, foxes, sables, martens, skunks, kolinskis, fitch, fishers, ermines, cats, sea otters, fur seals, hair seals, lions, tigers, leopards, lynxes, jackals, &c. The Rodentia include beavers, nutrias, musk-rats or musquash, marmots, hamsters, chinchillas, hares, rabbits, squirrels, &c. The Ungulata include Persian, Astrachan, Crimean, Chinese and Tibet lambs, mouflon, guanaco, goats, ponies, &c. The Marsupialia include opossums, wallabies and kangaroos.
Arsinoitherium is the precursor of the horned Ungulata; while Moeritherium and Palaeomastodon undoubtedly include the oldest known elephants.
ANCYLOPODA, or Ancylodactyla, an apparently primitive extinct subordinal group of Ungulata showing certain resemblances to the Perissodactyla, both as regards the cheek-teeth and the skeleton, but broadly distinguished by the feet being of an edentate type, carrying long curved and cleft terminal claws.
Of ungulata, besides a few hundreds of rare varieties, there are the springbuck, of which great herds still wander on the open veld, the steinbok, a small and beautiful animal which is sometimes coursed like a hare, the klipspringer or " chamois of South Africa," common in the mountains, the wart-hog and the dassie or rock rabbit.
Ungulata (Hoofed Mammals) :- a.
Whether the extinct Tillodontia are most nearly allied to the Rodentia, the Carnivora or the Ungulata, and whether they are really entitled to constitute an ordinal group by themselves, must remain for the present open questions.
An analogous statement may be made with regard to the sea-cows, or Sirenia, which appear to be derivates from the great herbivorous order of Ungulata, and might consequently be included in that group, as indeed has been already done in Dr Max Weber's classification.
It is with the proboscidean suborder of the Ungulata to which the Sirenia are most nearly related; the nature of this relationship being described by Dr Andrews as follows: " In the first place, the occurrence of the most primitive Sirenians with which we are acquainted in the same region as the most generalized proboscidean, Moeritherium, is in favour of such a view, and this is further supported by the similarity of the brain-structure and, to some extent, of the pelvis in the earliest-known members of the two groups.
In the case of the great order, or assemblage, of Ungulata it is necessary to pay somewhat more attention to fossil forms, since a considerable number of groups are either altogether extinct or largely on the wane.