Deep to these is the ovarian stroma, composed of fibrous tissue, and embedded in it are numerous nests of epithelial cells, the Graafian follicles, in various stages of development.
The fundamental substance or stroma is colorless and homogeneous.
Other series of modifications arise in which the tissues corresponding to the stroma invest the sporogenous hyphal ends, and thus enclose the spores, asci, basidia, &c., in a cavity.
Another simple case is where the plane or slightly convex surface of the stroma rises at its margins and overgrows the sporogenous hyphal ends, so that the spores, asci, &c., come to lie in the depression of a cavity - e.g.
Here we have the cushion-like type (stroma) of Nectria and many Pyrenomycetes, the clavate "receptacle" of Clavaria, &c., passing into the complex forms met with in Sparassis, Xylaria, Polyporei, and Agaricini, &c. In these cases the compound sporophore is often termed the hymenophore, and its various parts demand special names (pileus, stipes, gills, po--es, &c.) to denote peculiarities of distribution of the hymenium owlthe surface.
In other cases (Hypomyces, Nectria) the perithecia arise on an already mature stroma, while yet more numerous examples can be given (Poronia, Hypoxylon, Claviceps, &c.) where the perithecia originate below the surface of a stroma formed long before.
The simpler forms bear the perithecia directly on the mycelium, but the more highly developed forms often bear them on a special mycelial development - the stroma, which is often of large size and special shape and colour, and of dense consistence.
(Orlando Bandinelli), pope from 1159 to 1181, was a Siennese, and as a teacher of canon law in Bologna composed the Stroma or the Summa Magistri Rolandi, one of the earliest commentaries on the Decretum Gratiani.
Such characters are found in any primitive, sedentary group. More peculiarly Echinoderm features, in which the Pelmatozoan nature is manifest, are the enclosing of the viscera in a calcified and plated theca, for protection against those enemies from which a fixed animal cannot flee; the development, at the aboral pole of this theca, of a motor nerve-centre giving off branches to the stroma connecting the various plates of the theca and of its brachial, anal, and columnar extensions, and thus coordinating the movements of the whole skeleton; the absence of suckers from the podia, which, when present, are respiratory, not locomotor, in function.
Originally the calcareous substance of the plates (stereom) was pierced by irregular canals, more or less vertical, and containing strands of the soft tissue (stroma) that deposited the stereom, as well as spaces filled with fluid.
Io) the canals, that is to say, the stroma-strands, came to lie parallel to the surface and to cross the sutures between the plates, which were thus more flexibly and more strongly united; since the canals crossing each suture naturally occupy a rhombic area, the order is called Rhombifera.
Regarding the Echinoderms as a whole in the light of the foregoing account, we may give the following analytic summary of the characters that distinguish them from other coelomate animals: They live in salt or brackish water; a primitive bilateral symmetry is still manifest in the right and left divisions of the coelom; the middle coelomic cavities are primitively transformed into two hydrocoels communicating with the exterior indirectly through a duct or ducts of the anterior coelom; stereom, composed of crystalline carbonate of lime, is, with few exceptions, deposited by special amoebocytes in the meshes of a mesodermal stroma, chiefly in the integument; reproductive cells are derived from the endothelium, apparently of the anterior coelom; total segmentation of the ovum produces a coeloblastula and gastrula by invagination; mesenchyme is formed in the segmentation cavity by migration of cells, chiefly from the hypoblast.
The stereom and stroma become arranged in folds and strands at right angles to the sutures of the thecal plates; in higher forms the stereom-folds are in part specialized as pectini-rhombs.
An aborally-placed motor nerve-centre gives off branches to the stroma connecting the various plates of the theca and of its brachial, anal and columnar extensions, and thus co-ordinates the movements of the whole skeleton.