Lanistes among Streptoneura, Limacinidae among Opisthobranchia).
The fact that all the Euthyneura are hermaphrodite is not a fundamental difference; several Streptoneura are so, likewise Valvata, Oncidiopsis, Marsenina, Odostomia, Bathysciadium, Entoconcha).
- Streptoneura In this division the torsion of the visceral mass and visceral commissure is at its maximum, the latter being twisted into a figure of eight.
At the same time, the junction of the visceral loop above the intestine prevents in all Streptoneura the shortening of the visceral loop, and it is rare to find a fusion of the visceral ganglia with either pleural, pedal or cerebral - a fusion which can and does a, Anus.
The sexes are distinct, as in all Streptoneura; and genital ducts and accessory glands and pouches are present, as in all Pectinibranchia.
Visceral nerve-loop, the strands of which cross one another - this being characteristic of Streptoneura (Spengel).
In several of the more primitive forms the same torsion occurs as in Streptoneura, viz.
But in the other members of the sub-class the detorsion of the visceral mass has carried back the anus and circumanal complex from the anterior dorsal region to the right side, as in Bulla and Aplysia, or even to the posterior end of the body, as in Philine, Oncidium, Doris, &c. Different degrees of the same process of detorsion are, as we have seen, exhibited by the Heteropoda among the Streptoneura, and both in them and in the Euthyneura the detorsion is associated with degeneration of the shell.
The members of the order present the same wide range of superficial appearance as do the Pectinibranchiate Streptoneura, forms carrying well-developed spiral shells and large mantle-skirts being included in the group, together with flattened or cylindrical slug-like forms. But in respect of the substitution of other parts for the mantle-skirt and for the gill which the more degenerate Opisthobranchia exhibit, this order stands alone.
The torsion of the visceral hump is not carried out very fully, the consequence being that the anus has a posterior position a little to the right of the median line above the metapodium, whilst the branchial chamber formed by the overhanging mantle-skirt faces the right side of the body instead of lying well to the front as in Streptoneura and as in Pulmonate Euthyneura.
This enclosure of the shell is a permanent development of the arrangement seen in many Streptoneura (e.g.
Sp, Abdominal ganglion which represents also the supra-intestinal ganglion of Streptoneura and gives off the nerve to the osphradium (olfactory organ) o, and another to an unlettered socalled " genital " ganglion.
An operculum is present only in Amphibola; a contrast being thus afforded with the operculate pulmonate Streptoneura (Cyclostoma, &c.), which differ in other essential features of structure from the Pulmonata.
In some Pulmonata (snails) the foot is extended at right angles to the visceral hump, which rises from it in the form of a coil as in Streptoneura; in others the visceral hump is not elevated, but is extended with the foot, and the shell is small or absent (slugs).
Pulmonata are widely distinguished from a small number of Streptoneura at one time associated with them on account of their mantle-chamber being converted, as in Pulmonata, into a lung, and the ctenidium or branchial plume aborted.
The terrestrial Streptoneura (represented in England by the common genus Cyclostoma) FIG.
In the marine Streptoneura they are ectodermic projections which ultimately fall off; in the Opisthobranchs they are closed pouches; in Paludina and Bithynia they are canals as in Pulmonata.