I3.Typical siphonostele of dicotyledn.
To this type of steIn having a ground-tissue pith, whether with or without internal phloem, is given the name siphonostele to distinguish it from the solid haplostele characteristic of the root, the first-formed portion of the stem, and in the more primitive Pteridophytes, of the whole of the axis.
The type of siphonostele characteristic of many ferns, in which are found internal phloem, and an internal endodermis separating the vascular conjunctive from the pith is known as a solenostele.
Among the Cycadofilices a series of stages is found leading from the primitive fern-protostele to the type of siphonostele characteristic of the Cycads which agrees in essentials in all the Spermophytes.
The young tissue of the stelar cylinder, in the case of the modified siphonostele characteristic of the dicotyledonous stem, differs from the adjoining pith and cortex in its narrow elongated cells, a difference produced by the stopping of transverse and the increased frequency of longitudinal divisions.