The four sepals are arranged symmetrically, an outer median and an inner lateral pair.
From the upper rim of the receptacle are given off the five sepals, the five petals, and the very numerous stamens.
They are distinguished by having one of the five blue or yellow coloured sepals (the posterior one) in the form of a helmet; hence the English name monkshood.
There are five free sepals, overlapping in the bud, and, alternating with these, five free petals.
An outer whorl of five opposite the petals alternates with an inner whorl of five opposite the sepals; at the base of each of the antisepalous stamens is a honey-gland.
The sepals, very rarely three, which are two in number, fall off as the flower opens, the four (very rarely five or six) petals, which are crumpled in the bud stage, also fall readily.
The Californian poppy (Platystemon californicus) is a pretty annual about a foot high, having yellow flowers with 3 sepals and 6 petals; and the white bush poppy (Romneya Coulteri) is a very attractive perennial and semishrubby plant 2-8 ft.
In the second case the outer series (calyx of sepals) is generally green and leaf-like, its function being to protect the rest of the flower, especially in the bud; while the inner series (corolla of petals) is generally white or brightly coloured, and more delicate in structure, its function being to attract the particular insect or bird by agency of which pollination is effected.
The sepals are leafy and persistent; the corolla is generally divided into a longer or shorter tube and a limb which is spreading, as in primrose, or reflexed, as in Cyclamen; in Soldanella it is bell-shaped; in Lysimachia the tube is often very short, the petals appearing almost free; in Glaux the petals are absent.
Opposite to its lobes; this anomalous position is generally explained by assuming that an outer whorl of stamens opposite the sepals has disappeared, though sometimes represented by scales as in Samolus and Soldanella.
Another explanation is based on the late appearance of the petals in the floral development and their origin from the backs of the primordia of the stamens; it is then assumed that three alternating whorls only are present, namely, sepals, stamens bearing petal-like dorsal outgrowths, and carpels.
In diameter, with five spreading white petals alternating with five persistent green sepals, a large number of stamens with pinkish-brown anthers, and one to three carpels sunk in the cup-shaped floral axis.
The irregular flowers have five sepals united at the base, the dorsal one produced into a spurred development of the axis; of the five petals the two upper are slightly different and stand rather apart from the lower three; the eight stamens are unequal and the pistil consists of three carpels which form a fleshy fruit separating into three one-seeded portions.
The flowers are regular and symmetrical, having five sepals, tapering to a point and hairy on the margin, five petals which speedily fall, ten stamens, and a pistil bearing five distinct styles.
Arose from the supposed resemblance of the corona to the crown of thorns, and of the other parts of the flower to the nails, or wounds, while the five sepals and five petals were taken to symbolize the ten apostles - Peter, who denied, and Judas, who betrayed, being left out of the reckoning.
2) consists of a receptacle varying in form from that of a shallow saucer to that of a long cylindrical or trumpet-shaped tube, thin or fleshy in consistence, and giving off from its upper border the five sepals, the five petals (rarely these latter are absent), and the threads or membranous processes constituting the "corona."
The flowers are regular and rather showy, generally with three greenish sepals, followed in regular succession by three white or purplish petals, six to indefinite stamens and six to indefinite free carpels.
2), which is also dioecious, the small male flowers are borne in large numbers in shortstalked spathes; the petals are minute and scalelike, and only two of the three stamens are fertile; the flowers become detached before opening and rise to the surface, where the sepals expand and form a float bearing the two projecting semi-erect stamens.
Sepals transformed into leaves.
22, we recognize four distinct whorls of leaves: an outer whorl, the calyx of sepals; within it, another whorl, the parts alternating with those of the outer whorl, the corolla of petals; next a whorl of parts alternating with the parts of the corolla, the androecium of stamens; and in the centre the gynoecium of carpels.
The sepals are generally of a greenish colour; their function is mainly protective, shielding the more delicate internal organs before the flower opens.
In some cases the petals have the appearance of sepals, then they are sepaloid, as in Juncaceae.
S, Sepals; p, petals; a, stamens; c, carpels.
S, Sepals; p, petals; a, stamens; c, carpels.
In some cases, as in the vine-family Ampelidaceae, this seems to be the ordinary mode of development, but the superposition of the stamens on the sepals in many plants, as in the pink family, Caryophyllaceae, is due to the suppression or abortion of the whorl of petals, and this idea is borne out by the development, in some plants of the order, of the suppressed whorl.
Thus, a symmetrical flower may have five sepals, five petals, five stamens and five carpels, or the number of any of these parts may be ten, twenty or some multiple of five.
33 is a diagram of a symmetrical flower of stone-crop, with five sepals, five alternating petals, ten stamens and five carpels.
- Diagrammatic section of a symmetrical pentamerous flower of Stone-crop (Sedum), consisting of five sepals (s), five petals (p) alternating with the sepals, ten stamens (a) in two rows, and five carpels (c) containing ovules.
- Diagram of the flower of Flax (Linum), consisting of five sepals (s), five petals (p), five stamens (a), and five carpels (c), each of which is partially divided into two.
Angles formed by the lateral and median planes are the diagonal planes and in these flowers the petals which alternate with the sepals are cut by the diagonal planes.
- Diagram of flower of Sweet-pea (Lathyrus), showing five sepals (s), two superior, one inferior, and two lateral; five petals (p), one superior, two inferior, and two lateral; ten stamens in two rows (a); and one carpel (c).
Thus, in many Caryophyllaceae, as Polycarpon and Holosteum, while the calyx and corolla are pentamerous, there are only three or four stamens and three carpels; in Impatiens Noli-me-tangere the calyx is composed of three parts, while the other verticils have five; in labiate flowers there are five parts of the calyx and corolla, and only four stamens; and in Tropaeolum pentaphyllum there are five sepals, two petals, eight stamens and three carpels.
The symmetry in the flower is evidently dimerous, and the abnormality in the androecium, where the four long stamens are opposite the posterior sepals, takes place by a splitting, at a very early stage of development, of a single outgrowth into two.
The following is a very simple mode which has been proposed: - The several whorls are represented by the letters S (sepals), P (petals), St (stamens), C (carpels), and a figure marked after each indicates the number of parts in that whorl.
As regards each leaf of the flower, it is either spread out, as the sepals in the bud of the lime-tree, or folded upon itself (conduplicate), as in the petals of some species of Lysimachia, or slightly folded inwards or outwards at the edges, as in the FIG.
The sepals are sometimes free or separate from each other, at other times they are united to a greater or less extent; in the former case, the calyx is polysepalous, in the latter gamosepalous or monosepalous.
Sepals are generally of a more or less oval, elliptical or oblong form, with their apices either blunt or acute.
In a polysepalous calyx the number of the parts is indicated by Greek numerals prefixed; thus, a calyx which has three sepals is trisepalous; one with five sepals is pentasepalous.
The sepals occasionally are of different forms and sizes.
In a gamosepalous calyx the sepals are united in various ways, sometimes very slightly, and their number is marked by the divisions at the apex.
There are two sepals which fall off before the petals expand.
The part formed by the union of the sepals is called the tube of the calyx; the portion where the sepals are free is the limb.
Occasionally, certain parts of the sepals undergo marked enlargement.
In the violet the calycine segments are prolonged downwards beyond their insertions, and in the Indian cress (Tropaeolum) this prolongation is in the form of a spur (calcar), formed by three sepals; in Delphinium it is formed by one.
In Pelargonium the spur from one of the sepals is adherent to the flower-stalk.
In Potentilla and allied genera an epicalyx is formed by the development of stipules from the sepals, which form an apparent outer calyx, the parts of which alternate with the true sepals.
In Eschscholtzia and Eucalyptus the sepals remain united at the upper part, and become disarticulated at the base or middle, so as to come off in the form of a lid or funnel.
Petals differ more from ordinary leaves than sepals do, and are much more nearly allied to the staminal whorl.
Longitudinal section of flower; v, bracteole on the peduncle; 1, sepals; ls, appendage of sepal; c, petals; cs, spur of the lower petals; fs, glandular appendage of the lower stamens; a, anthers.
Thus, in a phanerogam, the sepals, petals, stamens and foliage-leaves all come under the category leaf, though some are parts of the perianth, others are spore-bearing organs (sporophylls), and others carry on nutritive processes.
Accepting this view of the phylogeny of the leaf, the perianthleaves (sepals and petals) and the foliage-leaves may be regarded as modified or metamorphosed sporophylls; that is, as leaves which are adapted to functions other than the bearing of spores.
The sepals are generally organs for the protection of the flower-bud; the petals, for attracting insects by their conspicuous form and color; the foliage-leaves, for the assimilation of carbon dioxide and other associated functions.
The parts of the flower are most frequently arranged in fives, or multiples of fives; for instance, a common arrangement is as follows, - five sepals, succeeded by five petals, ten stamens in two sets of five, and five or fewer carpels; an arrangement in fours is less frequent, while the arrangement in threes, so common in monocotyledons, is rare in dicotyledons.
Very beautiful also are the red velvet or white velvet sepals of the Mussaenda genus.
Pleurothallidinae, characterized by a thin stem bearing one leaf which separates at a distinct joint; the sepals are usually much larger than the petals and lip. Includes To genera, natives of tropical America, one of which, Pleurothallis, contains about 400 species.
And Sambucus, more rarely two-lipped as in Lonicera; the sepals and petals are usually five in number and placed above the ovary, the five stamens are attached to the corolla-tube, there are three to five carpels, and the fruit is a berry as in honeysuckle or snowberry (Symphoricarpus), or a stone fruit, with several, usually three, stones, as in Sambucus.
The flowers have a hollow tube at the base bearing at its free edge five sepals, an equal number of petals, usually concave or spoon-shaped, pink or white, and a great number of stamens.
The flowers are regular, with four free sepals arranged in two pairs at right angles, four petals arranged crosswise in one series, and two sets of stamens, an outer with two members and an inner with four, in two pairs placed in the middle line of the flower and at right angles to the outer series.
The flowers, which are solitary, or rarely in pairs, at the end of slender axillary flower-stalks, are very irregular in form, with five sepals prolonged at the base, and five petals, the lowest one larger than the others and with a spur, in which collects the honey secreted by the spurs of the two adjoining stamens.
B, pair of bracteoles below the flower; s, sepals; p, petals; st, stamens; o, ovary.
The sepals, which are generally free, show much variation in size, shape and covering, and afford valuable characters for the distinction of genera or sub-genera.
The sepals and petals are free or more or less united, the stamens as many or twice as many as the petals; the carpels, usually free, are equal to the petals in number, and form in the fruit follicles with two or more seeds.