It is a top-shaped, free-swimming organism provided with a preoral band of cilia,.
The antennules (or first antennae) are almost universally regarded as true appendages, though they differ from all the other appendages in the fact that they are always innervated from the " brain " (or preoral ganglia), and that they are uniramous in the nauplius larva and in all the Entomostracan orders.
The antennae (second antennae) are of special interest on account of the clear evidence that, although preoral in position in all adult Crustacea, they were originally postoral appendages.
Thus the bilateral symmetry would have been accentuated, and the organism shaped more definitely into three segments, namely (r) a preoral segment or lobe, containing the anterior coelomic cavity; (2) a middle segment, containing the gut, and the two middle coelomic cavities; (3) a posterior segment, containing the posterior coelomic cavity, which, however, owing to the backward prolongation of the anus, became divided into two - a right and left posterior coelom.
The epithelium of the outer surface was probably ciliated, and a portion of it in the preoral lobe differentiated as a sense-organ, with longer cilia and underlying nerve-centre, from which two nerves ran back below the ventral surface.
The forward portion of the anterior coelom shared in the constriction and elongation of the preoral lobe; but its hinder portion was dragged up along with the water-pore and formed a canal lying moccth ive along the outer wall (the gerei tatpore parietal canal).
The chief modifications of this form are seen in the Mitraria larva of Ammochares with only the preoral band, which is much folded and which has provisional and long setae; the a.trochous larva, where the covering of cilia is uniform and not split into bands; and the polytrochous larva where there are several bands surrounding the body.
In development some Lamellibranchia pass through a free-swimming trochosphere stage with preoral ciliated band; other fresh (I) ' Av B m 10 ad 3r ?/ a.: a il J FIG.
Ment and of the struc ture of the disk to their identification of "trochus" and "cingulum" with the preoral and postoral wreaths of the trochophore larva.