Sharp (1899), who divides the order into six great series of families: Lamellicornia (including the chafers and stag-beetles and their allies with five-segmented feet and plate-like terminal segments to the feelers); Adephaga (carnivorous, terrestrial and aquatic beetles, all with five foot-segments); Polymorpha (including a heterogeneous assembly of families that cannot be fitted into any of the other groups); Heteromera (beetles with the fore and intermediate feet five-segmented, and the hind-feet four-segmented); Phytophaga (including the leaf-beetles, and longhorns, distinguished by the apparently four-segmented feet), and Rhynchophora (the weevils and their allies, with head prolonged into a snout, and feet with four segments).
Lameere (1900) has suggested three sub-orders, the Cantharidif ormia (including the Phytophaga, the Heteromera, the Rhynchophora and most of the Polymorpha of Sharp's classification), the Staphyliniformia (including the rove-beetles, carrion-beetles and a few allied' amilies of Sharp's Polymorpha), and the Carabidiformia (Adephaga).
Kolbe (1901), who recognizes three sub-orders: (i.) the Adephaga; (ii.) the Heterophaga, including the Staphylinoidea, the Actinorhabda (Lamellicornia), the Heterorhabda (most of Sharp's Polymorpha), and the Anchistopoda (the Phytophaga, with the ladybirds and some allied families which Sharp places among the Polymorpha); (iii.) the Rhynchophora.
In regions of heavy rainfall the ohia-lehua (Metrosideros polymorpha), a tree growing from 30 to 100 ft.
Mushketov [1850-1902 ]) Cardium edule, Dreissena polymorpha, Neritina liturata, Adacna vitrea, Hydrobia stagnalis, in the Kara-kum desert, and Lithoglyphus caspius, Hydrobia stagnalis, Anodonta ponderosa and the sponge Metchnikovia tuberculata, in the Kizilkum desert.