The phloem is generally Type.
Eventually the new phellogens reach the level of the secondary phloem, and are formed in the parenchyma of the latter, keeping pace in their inward march with the formation of fresh secondary phloem by the cambium.
The xylem and phloem also, rarely form perfectly continuous layers as they do in a solenostelic fern.
The cylinder is surrounded by a mantle of one or more layers of parenchymatous cells, the pericycle, and the xylem is generally separated from the phloem in the stem by a similar layer, the mesocycle (corresponding with the amylom sheath in mosses).
The Stele In But, unlike the ferns, there is in the seed-plants no in- s d I ~ ternal phloem (except as a special development in ee pan $~$~ certain families) and no internal endodermis.
When the pith is large celled, the xylems of the bundles are separated from it by a distinct layer of conjunctive tissue called the endocycle, and a similar layer, the pericycle, separates the phloem from the cortex.
The xylem and phloem parenchyma consist of living cells, fundamentally similar in most respects to the medullary ray cells, which sometimes replace them altogether.
A considerable evolution in complexity can be traced in passing from the simplest forms of xylem and phloem found in the primary vascular tissues both among Pteridophytes and Phanerogams to these highly differentiated types.
In the primary and secondary tissue, is that the proteid cells of the phloem are here always sister-cells of the leptoids and are known as companion-cells.
Sometimes in such cases the cambium ceases to be active round these bays and joins across the outside of the bay, where it resumes its normal activity, thus isolating a phtoem strand, or, as it is sometimes called, a phloem -island, in the midst of the xylem.
Opposite the primary xylems, the cambium either (a) forms parenchyma on both sides, making a broad, secondary (principal) ray, which interrupts the vascular ring and is divided at its inner extremity by the islet of primary xylem; or (b) forms secondary xylem and phloem in the ordinary way, completing the vascular ring.
The water taken up by the root from the soil contains nitrogenous and mineral salts which combine with the first product of photo-synthesis - a carbohydrate - to form more complicated nitrogen-containing food substances of a proteid nature; these are then distributed by other elements of the vascular bundles (the phloem) through the leaf to the stem and so throughout the plant to wherever growth or development is going on.
This phenomenon follows injury to the phloem in the lower parts of the stem, preventing the downward flow of elaborated sap. The injury may be due to gnawing insects, and particularly to the fungus Corticium vagum, var.
Phloem centripetal in its development.
The xylem and phloem are nearly always found in close association in strands of various shapes in all the three main organs of the sporophyteroot, stem and leafand form a connected tissue-system running through the whole body.
This consists of a few xylem elements, e a a segment of phloem, pericycle, and usually an arc of h~s endodermis, which closes round the bundle as it detaches ~
The latter is often sclerized, especially opposite the phloem, and to a less extent opposite the xylem, as in the stem.
As a bundle is traced towards its blind termination in the mesophyll the peridesmic stereom first disappears, the sieve-tubes of the phloem are replaced by narrow elongated parenchyma cells, which soon die out, and the bundle ends with a strand of tracheids covered by the phloeotermic sheath.
Differentiation of the xylem progresses outwards, of the phloem inwards, but the two tissues never meet in the centre.
The protoxylems and the phloem strands are developed alternately, just within the outer limit of the young cylinder.
The cambium in the root, which is found generally in those plants which possess a cambium in the stem, always begins in the conjunctive tissue internal to the primary phloems, and Camblum forms new (secondary) phloem in contact with the In Roots primary, and secondary xylem internally.
Secondary xylem and phloem produced by a single cambium, or by successive cambial zones; no true vessels (except in the Gnetales) in the wood, and no companioncells in the phloem.
The type of siphonostele characteristic of many ferns, in which are found internal phloem, and an internal endodermis separating the vascular conjunctive from the pith is known as a solenostele.
Haplostele, the segments of inner endodermis, pericycle, phloem and ~ Pig.
Strengthening tissue of all kinds (and sometimes even the phloem) is more or less rudimentary.
In the other groups of Pteridophytes internal phloem is not found and an internal endodermis but rarely.
The leaf-bundles are always collateral (the phloem being turned downwards and the xylem upwards), even in Ferns, where the petiolar strands are concentric, and they have the ordinary mesodesm and peridesm of the collateral bundle.
The sieve-tubes of the secondary phloem usually have very oblique end-walls bearing a row of sieve-plates; plates also occur on the radial side-walls.
In others the secondary phloem is produced more abundantly in those places where the secondary xylem is deficient, so that the stem remains cylindrical in section, the phloem occupying the bays left in the xylem mass.
The significance of these phenomena, which present many minor modifications in different cases, is nol fully understood, but one purpose of the formation of phloem promontories and islands seems to be the protection of the sieve-tubes from crushing by the often considerable peripheral pressure that is e~ercised on the stems of these lianes.
An ordinary cambium is scarcely ever found in the Monocotyledons, but in certain woody forms a secondary meristem is formed outside the primary bundles, and gives rise externally to a little secondary cortex, and internally to a secondary parenchyma in which are developed numerous zones of additional bundles, usually of concentric structure, with phloem surrounded by xylem.