The cylinder is surrounded by a mantle of one or more layers of parenchymatous cells, the pericycle, and the xylem is generally separated from the phloem in the stem by a similar layer, the mesocycle (corresponding with the amylom sheath in mosses).
Such a vascular cylinder is called a haplostele, and the axis containing it is said to be haplostelic. In the stele of the root the strands of tracheids along the lines where the xylem touches the pericycle are spiral or annular, and are the xylem elements first formed when the cylinder is developing.
Or many protoxylems. When the protoxylem strands are situated at the periphery of the stele, abutting on the pericycle, as in all roots, and many of the more primitive Pteridophyte stems, the stele is said to be exarch.
This consists of a few xylem elements, e a a segment of phloem, pericycle, and usually an arc of h~s endodermis, which closes round the bundle as it detaches ~
Where internal phloem is present this is separated from the internal endodermis by an endocycle or internal pericycle, as it is sometimes called, and from the xylem by an internal mesocyclethese two layers, together with the outer mesocycle and pericycle, constituting the conjunctive tissue of the now hollow cylindrical stele.
Haplostele, the segments of inner endodermis, pericycle, phloem and ~ Pig.
The pericycle and mesocycle together form the conjunctive tissue of the stele in these simplest types.
The pericycle, medullary rays, endocycle and mesoderm all form parts of one tissue system, the external conjunctive, and are only topographically separable.
The relatively peripheral position ii the stem of the pericycle is important in this connexmon.
The protoxylem and protophloem are developed a few cells from the inner and outer margins respectively of the desmogen strand, the desmogenic tissue left over giving rise to the segments of endocycle and pericycle capping the bundle.
The differentiation of the stelar stereom, which usually takes the form of a sclerized pericycle, and may extend to the endocycle and parts of the rays, takes place in most cases later than the formation of the primary vascular strand.
This is known as exogenous branch-formation In the root, on the other hand, the origin of branches is endogenous The cells of the pericycle, usually opposite a protoxylem strand divide tangentially and give rise to a new growing-point.
The connections of its stele witl that of the parent axis are made across the pericycle of the latter Its cortex is never in connection with the cortex of the parent, but with its pericycle.
The formation of additional cambial cylinders or bands occurs in the most various families of Dicotyledons and in some Gymnosperms. They may arise in the pericycle or endocycle of the stele, in the cortex of the stem, or in the parenchyma of the secondary xylem or phloem.
After the cambium has been active for some time producing secondary xylem and phloem, the latter consisting of sievetubes, phloem-parenchyma and frequently thick-walled fibres, a second cambium is developed in the pericycle; this produces a second vascular zone, which is in turn followed by a third cambium, and so on, until several hollow cylinders are developed.
The root is diarch in structure, but additional protoxylem-strands may be present at the base of the main root; the pericycle consists of several layers of cells.
A pine needle grown iji continuous light differs from one grown under ordinary conditions in the absence of hypodermal fibres, in the absence of the characteristic infoldings of the mesophyll cell-walls, in the smaller size of the resin-canals, &c. The endodermis in Pinus, Picea and many other genera is usually a well-defined layer of cells enclosing the vascular bundles, and separated from them by a tissue consisting in part of ordinary parenchyma and to some extent of isodiametric tracheids; but this tissue, usually spoken of as the pericycle, is in direct continuity with other stem-tissues as well as the pericycle.
It has been suggested that transfusion-tracheids represent, in part at least, the centripetal xylem, which forms a distinctive feature of cycadean leaf-bundles; these short tracheids form conspicuous groups laterally attached to the veins in Cunninghamia, abundantly represented in a similar position in the leaves of Sequoia, and scattered through the so-called pericycle in Pinus, Picea, &c. It is of interest to note the occurrence of precisely similar elements in the mesophyll of Lepidodendron leaves.
The climbing species of Gnetum are characterized by the production of several concentric cylinders of secondary wood and bast, the additional cambium-rings being products of the pericycle, as in Cycas and Macrozamia.
The anatomy of Lycopodium presents considerable variety in detail, but the stem is always monostelic and the development of the xylem centripetal, the protoxylems being situated at the periphery of the stele; pericycle and endodermis surround the stele, and the wide cortex may be more or less sclerenchymatous.
The endodermis and pericycle surround the whole stele in Botrychium and Helminthostachys; in Ophioglossum each bundle has a separate sheath.
Transverse section of stem, showing the pith containing groups of sclerotic cells, the primary xylem-strands, secondary wood and phloem, pericycle and cortex.
Adventitious roots, arising from stems, usuall) take origin in the pericycle, but sometimes from other parts of th Conjunctive.
When the pith is large celled, the xylems of the bundles are separated from it by a distinct layer of conjunctive tissue called the endocycle, and a similar layer, the pericycle, separates the phloem from the cortex.