- Wing-Neuration in a Cossid Moth.
Wings with predominantly longitudinal neuration, covered with flattened scales.
Apart from this feature the Trichoptera also differ from the typical Neuroptera in the relatively simple, mostly longitudinal neuration of the wings, the absence or obsolescence of the mandibles and the semi-haustellate nature of the rest of the mouth-parts.
In the form of the feelers, the wing-neuration and minor structural details there is much diversity among the saw-flies.
Needham the wing-neuration in this order is the most primitive to be found in the Hexapoda.
When wings are present the front pair are much larger than the hind pair, and the neuration is remarkable for the concresence of the median with the After Marlatt, Bull.
On the other hand, the reduced feelers, the numerous Malpighian tubes (40), the large complex eyes, the vestigial condition of the jaws, the excessive size of the fore-wings as compared with the hind-wings and their complex neuration with an enormous number of crossnervules are all specializations.
All the wings are of firm, glassy texture, and very complex in their neuration; a remarkable and unique feature is that a branch of the radius (the radial sector) crosses the median nervure, while, by the development of multitudinous cross-nervules, the wing-area becomes divided into an immense number of small areolets.
They may be distinguished from the Neuroptera by the elongation of the head into a beak, the small prothorax, the narrow, elongate wings with predominantly longitudinal neuration, the presence of abdominal cerci and the cruciform larva.
At the base of each wing projects a dorsal lobe - the jugumand the neuration is predominantly longitudinal, resembling so closely that of the lower Lepidoptera (q.v.) that a nearer relationship of the Trichoptera to that order than to any group of the old Linnean Neuroptera is certain.