The tergites of this region and those of the following region, the metasoma, are fused to form a second or posterior carapace in Limulus, whilst remaining free in Scorpio.
Passing on now from the mesosoma we come in Scorpio to the metasoma of six segments, the first of which is broad whilst the rest are cylindrical.
The somites of the metasoma carry no parapodia.
In Limulus the metasoma is practically suppressed.
42) the six somites of the mesosoma are not fused to form a carapace at an early stage, and they are followed by three separately marked metasomatic somites; the other three somites of the metasoma have disappeared in Limulus, but are represented (From Lankester, loc. cit.) by the unsegmented prae-anal region.
It is probable that we have in the metasoma of Limulus a case of the disappearance of once clearly demarcated somites.
Following the metasoma in Limulus, we have as in Scorpio the post-anal spine - in this case not a sting, but a powerful and important organ of locomotion, serving to turn the animal over when it has fallen upon its back.
Perhaps the most important general agreement of Scorpio compared with Limulus and the Eurypterines is the division of the body into the three regions (or tagmata) - prosoma, mesosoma and metasoma - each consisting of six segments, the prosoma having leg-like appendages, the mesosoma having foliaceous appendages, and the metasoma being destitute of appendages.
The unsegmented dense chitinous sternal plate of the metasoma (XIII to XVIII) is not removed.
We shall, therefore, ignoring the ocular somite, speak of the first, second, third, fourth, fifth and sixth legbearing somites of the prosoma, and indicate the appendages by the Roman numerals, I, II, III, IV, V, VI, and whilst ignoring the praegenital somite we shall speak of the first, second, third, &c., somite of the mesosoma or opisthosoma (united mesosoma and metasoma) and indicate them by the Arabic numerals.
Intermediate somites forming a mesosoma occur, but tend to fuse superficially with the metasomatic carapace or to become co-ordinated with the somites of the metasoma, whether fused or distinct to form one region, the opisthosoma (abdomen of authors).
In the most highly developed forms the two anterior divisions (tagmata) of the body, prosoma and mesosoma, each exhibit six pairs of limbs, pediform and plate-like respectively, whilst the metasoma consists of six limbless somites and a post-anal spine.
(After Beecher.) ancestor of the grade) grouped in three regions of six - (a) the " prosoma " with palpiform appendages, (b) the " mesosoma " with plate-like appendages, and (c) the " metasoma " with suppressed FIG.
Concentration of the organ-systems by fusion of neighbouring regions (prosoma, mesosoma, metasoma), pre viously distinct, has frequently occurred, together with obliteration of the muscular and chitinous structures indicative of distinct somites.
- Nomomeristic Arachnids, in which the somites corresponding to mesosoma and metasoma have entirely aborted.
- These start from highly developed and specialized aquatic branchiferous forms, exhibiting a prosoma with six pediform pairs of appendages, an intermediate prae-genital somite, a mesosoma of six somites bearing lamelliform pairs of appendages, and a metasoma of six somites devoid of appendages, and the last provided with a post-anal spine.
Not freeswimming, none of the prosomatic appendages modified to act as paddles; segments of the mesosoma and metasoma (= opisthosoma) not more than ten in number, distinct or coalesced.
- Free-swimming forms, with the appendages of the 6th or 5th and 6th pairs flattened or lengthened to act as oars; segments of mesosoma and metasoma (= opisthosoma), twelve in number.
- The primitive distinction between the mesosoma and the metasoma retained, the latter consisting of six somites and the former of six somites in the adult, each of which is furnished during growth with a pair of appendages.
- The primitive distinction between the mesosoma and the metasoma wholly or almost wholly obliterated, the two regions uniting to form an opisthosoma, which never consists of more than twelve somites and never bears appendages or breathing-organs behind the 4th somite.