A leaf-trace, as it passes through the cortex, has a collateral structure, the protoxylem being situated at the inner edge of the xylem; when it reaches the leaf-base the position of the spiral tracheids is gradually altered, and the endarch arrangement (protoxylem internal) gives place to a mesarch structure (protoxylem more or less central and not on the edge of the xylem strand).
Similarly in the sporophylls of some cycads the bundles are endarch near the base and mesarch near the distal end of the stamen or carpel.
The mesarch structure of the leaf-bundles is met with in a less pronounced form in the flower peduncles of some cycads.
Palaeozoic genera, has not entirely disappeared from the stems of modern cycads; but the mesarch bundle is now confined to the leaves and peduncles.
A point of anatomical interest is the occurrence in the vascular bundles of the cotyledons, scale-leaves, and elsewhere of a few centripetally developed tracheids, which give to the xylem-strands a mesarch structure such as characterizes the foliar bundles of cycads.
The leaf-traces, where they traverse the cortex, have the structure of the foliar bundles in Cycads, for they are of the collateral type, and their xylem is mesarch, the spiral elements lying in the interior of the ligneous strand.