The leaves of the true mosses and those of the club-mosses (Lycopodium, Selaginella) being somewhat alike in general appearance and in ontogeny, might be, and indeed have been, regarded as homologous on that ground.
In his experiments upon this subject Fraunhofer employed plates of glass dusted over with lycopodium, or studded with small metallic disks of uniform size; and he found that the diameters of the rings were proportional to the length of the waves and inversely as the diameter of the disks.
In diameter, has a little lycopodium powder introduced, and the powder is allowed to run all along the tube, which is then fixed horizontally.
Stationary waves are formed in the air in the dust-tube if the length is rightly adjusted by the closely-fitting piston, and the lycopodium dust collects at the nodes in little heaps, the first being at the fixed end and the last just in front of the piston on the sounder.
If after the deposition of the drop, a little lycopodium be scattered over the surface, it is seen that a circular space surrounding the drop, of about the size of a shilling, remains bare, and this, however often the dusting be repeated, so long as any of the carbon bisulphide remains.
Small tuberous shoots, comparable on a large scale with the bulbils of Lycopodium Selago, are occasionally produced in the axils of some of the persistent leaf-bases; these are characteristic of sickly plants, and serve as a means of vegetative reproduction.
B, Lycopodium cernuum.
The sporangia of the Psilotaceae are associated in synangia, which occupy the same position relatively to the sporophyll, as the single sporangium of Lycopodium or the group of sporangia in Spenophyllum majus.
There is some reason to believe that the prothallus of Psilotum resmbles some Lycopodium prothalli, but conclusive evidence is wanting; that of Tmesipteris is unknown.
A number of species of Lycopodium are epiphytic and those of Isoetes live submerged in water.
The modifications shown by the gametophyte of Lycopodium will be described below.
This order contains the two genera Phylloglossum and Lycopodium; the former has a single species, confined to Australia, Tasmania and New Zealand, while nearly one hundred species of Lycopodium are known.
- Lycopodium clavatum.
The anatomy of Lycopodium presents considerable variety in detail, but the stem is always monostelic and the development of the xylem centripetal, the protoxylems being situated at the periphery of the stele; pericycle and endodermis surround the stele, and the wide cortex may be more or less sclerenchymatous.
The morphology of Stigmaria has been much discussed; possibly the main axes, which do not agree perfectly either with rhizomes or roots, may best be regarded as comparable with the rhizophores of Selaginellae; they have also been compared with the embryonic stem, or protocorm, of certain species of Lycopodium; the homologies of the appendages with the roots of recent Lycopods appear manifest.
This fact leads to the consideration of Phylloglossum, which resembles the embryo of Lycopodium cernuum in so many respects that it has been spoken of as a permanently embryonic form of Lycopod: it is in some respects the simplest existing Pteridophyte.
The sporangia agree with those of Lycopodium in structure and position.
The cones, which in some instances at least were heterosporous, presented a general resemblance to those of Lycopodium and Selaginella, a single sporangium being situated on the upper surface of each sporophyll.