Interradial tentacles may be also developed, so that the total number present may be increased to eight or to an indefinitely large number.
Preceded by six per cartilaginous margin of the radial and six interradial disk centripetally in the solid larval tentacles).
Outer surface of the jellyCartilaginous margin of the like disk; six of these are disk covered by thread perradial, six interradial, cells.
- Tropho- some unknown; gonosome, free medusae, with deep, bell-shaped umbrella, with interradial gonads on the base of the stomach, with branched radial canals, and correspondingly numerous hollow tentacles.
Primitively there are four perradial tentacles, to which may be added four interradial, or they may become very numerous and are then scattered evenly round the margin, never arranged in tufts or clusters.
The mesogloea becomes enormously increased in quantity (hence the popular name "jelly-fish"), and in correlation with this the endoderm-layer lining the coelenteron becomes pressed together in the interradial areas and undergoes concrescence, forming a more or less complicated gastrovascular system (see Medusa).
Usually the four subgenital cavities are distinct from each other (so-called tetrademnic condition), but in many Rhizostomeae, for example, Crambessa, the subgenital cavities join together under the subumbral floor of the stomach (so-called monodemnic condition) and coalesce to form a so-called subgenital portico placed on the oral side of the stomach, opening by four interradial apertures between the oral arms, that is to say, by the four primitive apertures of the subgenital pits.
The eight adradial gonads on the subumbral walls of the four radial pouches, representing primitively four horse-shoeshaped gonads each divided into two by an interradial septum.
Perradial, four interradial and ov, Ovaries.
Perradial and interradial canals near consist of a main stem giving off branches, and both stem and branches reach to the marginal ring-canal, the main stem ending in one of the eight tentaculocysts, which are lodged in the notches between the lobes of the umbrellar margin.
Oral surface of later stage of scyphistoma of Aurelia, with commencement of four interradial tentacles.
The four gastric interradial ridges are seen through the mouth.
The sixteen tentacles of the scyphistoma disappear, and in the place of the four perradial and four interradial tentacles, the eight tentaculocysts of the adult are formed as outgrowths of the subumbral margin, independently of the tentacles of the scyphistoma (Friedemann).
The ephyra has a flat, disk-shaped body, with eight marginal lobes (four perradial, four interradial); a tentaculocyst is lodged in a deep notch at the apex of each lobe.
As the 1 The four primitive interradial cathammata disappear in the fully formed ephyra and become replaced by sixteen subradial concrescence-areas without any ostia or ring-canal at the margin.
E, Young ephyra just liberated, showing the eight bifurcate arms of the disk ' and the interradial single gastral filaments.
In, Interradial radius.
- Medusae more or less cubical in form, with four perradial rhopalia alternating with four interradial tentacles or groups of tentacles; oral arms short; stomach a wide cavity bearing four interradial groups of phacellae and giving off four broad perradial pouches completely separated from each other by four interradial septa (i.e.
- With four interradial tentacles.
With four interradial groups of tentacles.
- With four interradial groups of tentacles, three in each group. Tripedalia.
Are interradial in position.
Semon called this stage the Pentactula, and supposed that, in its early history, the class had passed through a similar stage, which he called the Pentactaea, and regarded as the ancestor of all Echinoderms. It has since been proved that the five tentacles with their canals are interradial, so that one can scarcely look on the Pentactula as a primitive stage, while the apparent simplicity of the Synaptidae, at least as compared with other holothurians, is now believed to be the result of regressive vlu.
T, The five interradial tentacles.
The genital organs in both Asteroidea and Echinoidea would retain the interradial position they first assumed in Edrioaster; and in Echinoidea their primitive temporary openings to the exterior were converted into definite pores, correlated with five interradially placed plates at the aboral pole.
- Radial symmetry affects the food-grooves, and by degrees the thecal plates connected therewith, but not the interradial thecal plates; probably also the nerves and ambulacral vessels, but not the gonads.
Always), radiate from a central peristome between five interradial deltoid plates, and are edged by alternating side-plates bearing brachioles, to which side-branches pass from the grooves.
- Blastoidea in which the thecal plates have assumed a definite number and position in 3 circlets, as follows: 3 basals, 2 large and I small; 5 radials, often forkshaped, forming a closed circlet; 5 deltoids, interradial in position, supported on the shoulders or processes of the radials, and often surrounding the peristome with their oral ends.
Interradial, and called "basals (BB)."
- Crinoidea in which the base consists of BB only, the aboral prolongations of the chambered organ being interradial; new columnals are introduced at the extreme proximal end of the stem.
The comparative anatomy of living forms, combined with the evolutionary hypothesis sketched above, suggests that the early holothurians possessed the following characters: subvective grooves entirely closed; 5 radial canals, proceeding from the water-ring, gave off branches furnished with ampullae to the podia on each side of them, the 10 anterior podia being changed into cylindrical tentacles; the transverse muscles of the body-wall formed a circular layer, probably interrupted at the radii (though Ludwig believes the contrary); longitudinal muscles as paired radial bands, without those special retractors for withdrawing the anterior part of the body which occur in many recent forms; a hydropore connected with the water-ring by a canal in the dorsal mesentery; a gonopore behind the hydropore connected by a single duct with a bunch of genital pouches on each side of the mesentery; gut dextrally coiled, with a simple blood-vascular system, and with an enlargement at the anus for respiration, this eventually producing branched caeca called "respiratory trees"; skeleton reduced to a ring of 5 radial and 5 interradial plates round the gullet, and small plates, with a hexagonally meshed network, dispersed through the integument.