These constitute a hypodermal layer, beneath which the chlorophyll cells of the parenchyma are densely packed together, and are elongated in a direction vertical to the surface of the leaf, forming the palisade tissue.
The presence of hypodermal fibres is another feature worthy of note, but the occurrence of these elements is too closely connected with external conditions to be of much systematic value.
The change that is required to transform Exopterygota into Endopterygota is merely that a cell of hypodeimis should proliferate inwards instead of outwards, or that a minute hypodermal evaginated bud should be forced to the interior of the body by the pressure of a contracted cuticle.
A pine needle grown iji continuous light differs from one grown under ordinary conditions in the absence of hypodermal fibres, in the absence of the characteristic infoldings of the mesophyll cell-walls, in the smaller size of the resin-canals, &c. The endodermis in Pinus, Picea and many other genera is usually a well-defined layer of cells enclosing the vascular bundles, and separated from them by a tissue consisting in part of ordinary parenchyma and to some extent of isodiametric tracheids; but this tissue, usually spoken of as the pericycle, is in direct continuity with other stem-tissues as well as the pericycle.
The cortex is often preserved; in certain cases it was strengthened by hypodermal strands of fibres, as in Equisetum.
We are therefore entitled to assume that the suppressed wings of Exopterygota tend to reappear; and, speaking of the past, we may say that if after a period of suppression the wings began to reappear as hypodermal buds while a more rigid pressure was exerted by the cuticle, the growth of the buds would necessarily be inwards, and we should have incipient endopterygotism.
Tissues of the Cordaitean stem are well preserved; the cortex possessed a system of hypodermal strands of fibres, comparable to those found in the Lyginodendreae.