In this way the medusa sinks from an independent per sonality to an organ of the polyp-colony, becoming a so-called medusoid gonophore, or bearer of the reproductive organs, and losing gradually all organs necessary for an independent existence, namely those of sense, locomotion and nutrition.
41, C), where, however, there is a further complication in the form of an adventitious envelope or ectotheca (ect.) split off from the gonophore as a protective covering, and not present in Cladocoryne.
It is evident that the outer envelope of the gonophore represents the ex-umbral ectoderm (ex.), and that the inner ectoderm lining the cavity represents the sub-umbral ectoderm of the free medusa.
A, A male gonophore still enclosed in its ecto theca.
B and C, Two views of a female gonophore after t, Tentacles.
If the bud, however, is destined to give rise not to a free medusa, but to a gonophore, the development is similar but becomes arrested at various points, according to the degree to which the gonophore is degenerate.
The question is one intimately connected with the view taken as to the nature and individuality of polyp, medusa and gonophore respectively.
The hydroid genus Lafoea is remarkable for producing gonothecae on the hydrorhiza, each containing a blastostyle which bears a single gonophore; this portion of the colony was formerly regarded as an independent parasitic hydroid, and was named Coppinia.
It is a re markable fact that all specimens of Limnocodium hitherto seen have been males; it may be inferred from this either that only one polypstock has been introduced into Europe, from which all the medusae seen hitherto have been budded, or perhaps that the female medusa is a sessile gonophore, as in Pennaria.
Physalia, general view, diagrammatic; B, cormidium of Physalia; D, palpon; T, palpacle; G, siphon; GP, gonopalpon; M d', male gonophore; M y, female gonophore, ultimately set free.