All the appendages are pediform and bi-ramose; all have a prominent gnathobase, and in all the exopodite carries a comb-like series of secondary processes.
Stc, Sterno-coxal process (gnathobase) of the chelae.
In all the Phyllopoda the number of endites is six, and the proximal one is more or less distinctly specialized as a gnathobase, working against its fellow of the opposite side in seizing food and transferring it to the mouth.
The conversion of the Arthropod's limb into a jaw, as a rule, is effected by the development of an endite near its base into a hard, chitinized, and often toothed gnathobase (see figs.
(4) Three of the rami of the primitive limb (endites 5 and 6, and exite I) specially developed as endopodite, exopodite, and epipodite - the first two often as firm and strongly chitinized, segmented, leg-like structures; the original axis or corm reduced to a basal piece, with or without a distinct gnathobase (endite i)- typical tri-ramose limb of higher Crustacea.
(8) The gnathobase becomes greatly enlarged and not separated by a joint from the corm; it acts as a hemignath or half jaw working against its fellow of the opposite side.
The endopodite may be retained as a small segmented palp at the side of the gnathobase or disappear (mandible of Crustacea, Chilopoda and Hexapods).
The isolated gnathobase often seen as " mandible " and the genital operculum).
Had a gnathobase or jaw process on every parapodium.
As many as six pairs of appendages following the mouth may have an enlarged gnathobase actually functional as a jaw or hemignath, but a ramus is well developed on each of these appendages either as a simple walking leg, a palp or a chela.
In the more primitive forms the appendage of every post-oral somite has a gnathobase and two rami; in higher specialized forms the gnathobases may be atrophied in every appendage, even in the first post-oral.
The ancestral stock was (as in the Arachnida) pantognathobasic, that is to say, had a gnathobase or jaw-process on the base of every post-oral appendage.
A gnathobase is developed (in the primitive stock) on every pair of post-oral appendages; two prosthomeres present, the second somite as well as the first having passed in front of the mouth, but only the second has appendages.
The original stock, like that of the last grade, has a gnathobase on every post-oral appendage, but three prosthomeres are now present, in consequence of the movement of the oral aperture from the third to the fourth somite.
On the whole the facts seem to be against this supposition, though we need not suppose that the gnathobase was very large or the rami undeveloped in the buccal parapodia which were destined to lose their mandibular features and pass in front of the mouth.