1, C, sm, m - jointed on to the submentum, while the galeae, laciniae and palps remain distinct.
In specialized biting insects, such as beetles (Coleo C ptera), the labium tends to become a hard transverse plate bearing the pair of palps, a median structure - known as the ligula - formed of the conjoined laciniae, and a pair of small rounded processes - the reduced galeae - often called the " paraglossae," a term better avoided since it has been applied also to the maxillulae of Aptera, entirety different structures.
Mandibles absent in imago, very exceptionally present in pupa; first maxillae nearly always without laciniae and often without palps, or only with vestigial palps, their galeae elongated and grooved inwardly so as to form a sucking trunk.
E, First maxilla; a, cardo; b, stipes; c, galea; d, lacinia; e, palp. B, Second maxillae (Labium); a, mentum; b, ligula (between the two galeae); c, c, palps.