As in all such introand e-versible organs, eversion of the Gastropod proboscis is effected by pressure communicated by the muscular body-wall to the liquid contents (blood) of the body-space, accompanied by the relaxation of the muscles which directly pull upon either the sides or the apex of the tubular organ.
Supposing the tube to be completely introverted and to commence its eversion, we then find that eversion may take place, either by a forward movement of the side of the tube near its attached base, as in the proboscis of the Nemertine worms, the pharynx of Chaetopods and the eye-tentacle of Gastropods, or by a forward movement of the inverted apex of the tube, as in the proboscis of the Rhabdocoel Planarians, and in that of Gastropods here under consideration.
It is clear that, if we start from the condition of full eversion of the tube and watch the process of introversion, we shall find that the pleurecbolic variety is introverted by the apex of the tube sinking inwards; it may be called acrembolic, whilst conversely the acrecbolic tubes are pleurembolic. Further, it is obvious enough that the process either of introversion or of eversion of the tube may be arrested at any point, by the development of fibres connecting the wall of the introverted tube with the wall of the body, or with an axial structure such as the oesophagus; on the other hand, the range of movement of the tubular introvert may be unlimited or complete.
B, The same, partially everted by eversion of the sides, as in the Nemertine proboscis and Gastropod eye-tentacle = pleurecbolic.
D, E, A similar simple introvert in course of eversion by the forward movement, not of its sides, but of its apex, as in the proboscidean Rhabdocoels =acrecbolic.
G, The acrecbolic snout of a proboscidiferous Gastropod, arrested short of complete eversion by the fibrous band b.
I, Partial eversion of H.
K, Complete eversion of H.
This aggressive character has a different aspect in several genera which are destitute of a central stylet, but in which the surface that is turned outwards upon eversion of the proboscis is largely pro- P. vided with nematocysts, sending the urticating rods of different sizes in all directions.
The cloaca is often very large; the nephridia and oviducts may open into it, and the eggs lodge there on their way outwards; they are thrown out, as are the faecal masses, by an eversion of the cloaca.
The introvert is not a simple one with complete range both in eversion and introversion, but is arrested in introversion by the fibrous bands at c, and similarly in eversion by the fibrous bands at b.