The primitive shell-sac or shell-gland is well marked at this stage, and the pharynx is seen as a new ingrowth (the stomodaeum), about to fuse with and open into the primitively invaginated arch-enteron (fig.
Arch-enteron or cavity lined by the enteric cell-layer or endo derm.
The invaginated cells (derived from the division of the four big cells) form the endoderm or arch-enteron; the outer cells are the ectoderm.
Ing gs, the bilobed arch sh, The primitive shell-sac or enteron or lateral vesicles shell-gland ° of invaginated endoderm, pi, The rectal peduncle or whichwill develop into liver.
The body-cavity and the muscular, fibrous and vascular tissues are traced partly to two symmetrically disposed " mesoblasts," which bud off from the invaginated arch-enteron, partly to cells derived from the ectoderm, which at a very early stage is connected by long processes with the invaginated endoderm.
The mass of the arch-enteron or invaginated endodermal sac has taken on a bilobed form, and its cells are swollen (gs and tge).
Not only is the coelom thus subdivided, but the enteron (gut, alimentary canal, digestive tube) itself shows indications of three main subsections in continuity with one another: - (I) proboscis-gut (Eicheldarm, stomochord, vide infra); (2) collar-gut (buccal cavity, throat); (3) truncal gut extending from the collar to the vent.
25), no part of the blastopore persists either as mouth or as anus, but the aperture closes - the pedicle of invagination, or narrow neck of the invaginated arch-enteron, becoming the intestine.
24, in optical median section, showing the invaginated cells hy which form the arch-enteron, and the mesoblastic cells me which are budded off from the surface of the mass hy, and apply themselves to the inner surface of the epiblastic cell-layer cp. C, The same embryo focused so as to show the mesoblastic cells which immediately underlie the outer cell-layer.
The enteron itself becomes bilobed and is joined by a new invagination, that of the mouth and stomodaeum.
In the Coelentera, whatever subsequent changes of shape the little sac may undergo as it grows up to be polyp or jelly-fish, the original arch-enteron remains as the one cavity pervading all regions of the body.
In common with all other Coelomata, the Mollusca have the mouth and first part of the alimentary canal which leads into the met-enteron formed by a special invagination of the outer layer of the primitive body-wall, not to be confounded with that which often, but not always, accompanies the antecedent formation of the archenteron; this invagination is termed the stomodaeum.
Similarly an anal aperture is formed in connexion with a special invagination which meets the hinder part of the met-enteron, and is termed the proctodaeum.
Ae, Arch-enteron or cavity lined by the enteric cell-layer or endoderm.
The careful study of the development of one Acoelous form and of certain Rhabdocoels has strengthened this hypothesis by showing that no definite enteron or gut is at first laid down, but that certain embryonic syncytial tracts become digestive tracts, others excretory, others again muscular.