- Vacuolated Endoderm Cells of cartilaginous consistence from the axis of the tentacle of a Medusa (Cunina).
Further, two distinct types of otocyst can be recognized in the Hydro medusae; that of the Leptolinae, in which the entire organ is ectodermal, concrement-cells and all, and the organ is not a tentaculocyst; and that of the Trachylinae, in which the organ is a tentaculocyst, and the concrement-cells are endodermal, derived from the endoderm of the modified tentacle, while the rest of the organ is ectodermal.
- Tentaculocyst (statorhabd) ous with the endoderm of Cunina solnzaris.
In the endoderm large concretions are formed (con.).
The process carrying the otolith outer side of a or concretion hk, formed by endoderm cells, is tentacle, two enclosed by an upgrowth forming the " vesicle," nerves run round which is not yet quite closed in at the top. the base of the (After Hertwig.) tentacle to it.
The endoderm of the medusa shows the same general types of structure as in the polyp, described above.
We can distinguish (I) digestive endoderm, in the stomach, often with special glandular elements; (2) circu-, latory endoderm, in the radial and ring canals; (3) supporting endoderm in the axes of the tentacles and in the endodermlamella; the latter is primitively a double layer of cells, produced by concrescence OC-- = w.?"
40); in Geryonia, however, it remains double, and the centripetal canals arise by parting of the two layers; (4) excretory endoderm, lining pores at the margin of the umbrella, occurring in certain Leptomedusae as socalled " marginal tubercles," opening, on the one hand, into the ring-canal and, on the other hand, to the exterior by " marginal funnels," which debouch into the sub-umbral cavity above the velum.
As has been described above, the endoderm may also contribute to the sense-organs, but such contributions are always of an accessory nature, for instance, concrement-cells in the otocysts, pigment in the ocelli, and never of sensory nature, sense-cells being Hydromedusae are of separate sexes, the only known exception being Amphogona apsteini, one of the Trachomedusae (Browne ).
The reproductive cells may be regarded as belonging primarily to neither ectoderm nor endoderm, though lodged in the ectoderm in all Hydromedusae.
Cordylus of el, Endoderm lamella.
En, Endoderm lining the enteric cavity.
E, Wandering endoderm cells of the gelatinous substance.
G, With spadix prolonged e.1, Endoderm-lamella.
Have become obliterated by coalescence of their walls, so that the entire endoderm of the umbrella is in the condition of the endodermlamella.
The endodermal spadix (sp) of the sporosac represents the endoderm of the manubrium; the ectodermal lining of the sporosac (ex.) represents the ex-umbral ectoderm of the medusa; and the intervening layers, together with the sub-umbral cavity, have disappeared.
Of both layers, ecto;?, s.c. ?? ?.; derm and endoderm, s.c. n s ?;, v N containing a prolonga.
The endoderm is shaded, the ectoderm left clear.
E.l, Endoderm lamella.
Form the radial canals (r.c.), ring-canal (c.c.), and endoderm-lamella (e.l., fig.
The tissues of the bud become differentiated into ectoderm and endoderm, and the endoderm of the bud becomes secondarily continuous with that of the parent, but no part of the parental endoderm contributes to the building up of the daughter-polyp. Lang regarded this method of budding as universal in polyps, a notion disproved by O.
Endoderm, containing the st, Stomach, which in H ac coelenteric cavity (cod), quires a secondary com while the outer layer munication with the diges furnishes the future ectotive cavity of the mother.
We may distinguish the following series of stages: (I) ovum; (2) cleavage, leading to formation of a blastula; (3) formation of an inner mass or parenchyma, the future endoderm, by immigration or delamination, leading to the so-called parenchymula-stage; (4) formation of an archenteric cavity, the future coelenteron, by a splitting of the internal parenchyma, and of a blastopore, the future mouth, by perforation at one pole, leading to the gastrula-stage; (5) the outgrowth of tentacles round the mouth (blastopore), leading to the actinula-stage; and (6) the actinula becomes the polyp or medusa in the manner described elsewhere (see articles Hydrozoa, POLYP and Medusa).
Apart from the weighty arguments which the development furnishes against the theories of Allman and Mechnikov, it may be pointed out that neither hypothesis gives a satisfactory explanation of a structure universally present in medusae of whatever class, namely the endoderm-lamella, discovered by the brothers O.
It would be necessary to regard this structure as a secondary extension of the endoderm in the tentacle-web, on Allman's theory, or between the outgrowths of the hydrorhiza, on Mechnikov's hypothesis.
The Hertwigs when they discovered the endoderm-lamella showed on morphological grounds that polyp and medusa are independent types, each produced by modification in different directions of a more primitive type represented in development by the actinulastage.
The tentacles are always solid, containing an axis of endoderm-cells resembling notochordal tissue or plantparenchyma, and are but moderately flexible.
It consists of a hollow tube, or tubes, of which the wall is made up of the two body-layers, ectoderm and endoderm, and the cavity is a continuation of the digestive cavities of the nutritive and other appendages, i.e.
In the wall of the sack is a double layer of endoderm, the space between which is a continuation of the coelenteron.
By coalescence of the endoderm-layers, the coelenteron may be reduced to vessels, usually eight in number, opening into a ring-sinus surrounding the pore.
Thus the disposition of the endoderm-cavities is roughly comparable to the gastrovascular system of a medusa.
- The fertilized ovum gives rise to a parenchymula, with solid endoderm, which is set free as a free-swimming planula larva, in the manner already described (see Hydrozoa).
The endoderm of the planula now acquires a cavity, and at the narrower pole a mouth is formed, giving rise to the primary siphon.
The ectoderm is indicated by close hatching, the endoderm by light hatching, the mesogloea by thick black lines, the horny skeleton of the pneumatophore and sail by dotting.
The invaginated cells (derived from the division of the four big cells) form the endoderm or arch-enteron; the outer cells are the ectoderm.
Ing gs, the bilobed arch sh, The primitive shell-sac or enteron or lateral vesicles shell-gland ° of invaginated endoderm, pi, The rectal peduncle or whichwill develop into liver.
It has usually been regarded as representing both endoderm and mesoderm, and the groove which usually leads to its formation has been compared to the abnormally elongated blastopore of a typical gastrula.
These two endoderm-rudiments embryonic membrane formed by delamination from the blastoderm, ultimately grow together and give rise to the epithelium of the midwhile in a few insects, including the wingless spring-tails, the emgut.
He finds that the endoderm arises may be readily distinguished, six of which subsequently enter into from an anterior and a posterior rudiment derived from the " endothe formation of the head, three going to the thorax and twelve to blast," that many of the cells of these rudiments wander into the the abdomen.
Size compared with the thorax or abdomen, but in the embryo it On the whole it seems likely that the endoderm is represented in forms a much larger portion of the body than it does in the adult.
17), is the mouth or oral piece; the second, explained by the presence of a " latent endoderm-group " in those the antennal segment; the third, the intercalary or prae-mandibular invaginations.
There is some evidence that in this group the ectoderm of the oesophagus is chiefly concerned with digestion, whereas the endoderm of the intestine is limited to the absorption of the soluble products.
On the ether hand, a survey of the facts of cellular embryology which were accumulated in regard to a variety of classes within a few years of Kovalevsky's work led to a generalization, independently arrived at by Haeckel and Lankester, to the effect that a lower grade of animals may be distinguished, the Protozoa or Plastidozoa, which consist either of single cells or colonies of equiformal cells, and a higher grade, the Metazoa or Enterozoa, in which the egg-cell by " cell division " gives rise to two layers of cells, the endoderm and the ectoderm, surrounding a primitive digestive chamber, the archenteron.
(3) The generative cells are produced in either the ectoderm or endoderm, and not in a third layer arising in the embryo, distinct from the two primary layers; in other words, there is no mesoderm or coelom.
In this stage the body is composed of two layers, ectoderm (d) externally, and endoderm (c) internally, surrounding a central cavity, the archenteron (b), which communicates with the exterior by a pore (a), the blastopore.
Ep, Ectoderm; hy, endoderm; al, enteric cavity.
Between the ectoderm and endoderm a gelatinous supporting layer, termed the mesogloea, makes its appearance.
T, tentacle; g.p, gastric pouch; r.c, radial canal not present in C and C'; c.c, circular or ring-canal; e.1, endoderm-lamella formed by concrescence.
Further, in the hydropolyp the digestive cavity either remains simple and undivided and circular in transverse section, or may show ridges projecting internally, which in this case are formed of endoderm alone, without any participation of the mesogloea.
In the anthopolyp, on the other hand, the digestive cavity is always subdivided by so-called mesenteries, in-growths of the endoderm containing vertical lamellae of mesogloea (see Anthozoa).