This mode has some disadvantages attending it; such sheets are difficult to handle; the crustaceous species are liable to have their surfaces rubbed; the foliaceous species become so compressed as to lose their characteristic appearance; and the spaces between the sheets caused by the thickness of the specimen permit the entrance of dust.
(c) The crustaceous thallus, which is the most common of all, forms a mere crust on the substratum, varying in thickness, and may be squamose (in Squamaria), radiate (in Placodium), areolate, granulose or pulverulent (in various Lecanorae and Lecideae).
Calcium oxalate is a very common substance, especially in crustaceous lichens; fatty oil in the form of drops or as an infiltration in the membrane is also common; it sometimes occurs in special cells and in extreme cases may represent 90% of the dry substance as in Verrucaria calciseda, Biatora immersa.
Another very distinct group with silvery foliage - the crustaceous group - contains some of our choicest Alpines.
The asexual organs in the case of Cutleria multifida arise on a crustaceous form, Aglaozonia reptans, formerly considered to be a distinct species.