The ovary consists of numerous carpels united together and free, or more or less embedded in the top of the flower-stalk.
Axillary or terminal spikes; they have four stamens, which bear at the back four small herbaceous petal-like structures, and four free carpels, which ripen to form four small green fleshy fruits, each containing one seed within a hard inner coat;.
The ANGIOSFERMS, which are much the larger class, derive their name from the fact that the carpel or carpels form a closed chamber, the ovary, in which the ovules are developedassociated with this is the development of a receptive or stigmatic surface on which the pollen grain is deposited.
The sporophylls (stamens and carpels) are generally associated with other leaves, known as the perianth, to form a flower; these subsidiary leaves are protective and attractive in function and their development is correlated with the transport of pollen by insect agency (see ANGI0sPERM5; POLLINATION, and FLOWER).
Under the head of malformations we place cases of atrophy of parts or general dwarfing, due to starvation, the attacks of Fungi or minute insects, the presence of unsuitable food-materials and so on, as well as cases of transformation of stamens into petals, carpels into leaves, and so forth.
The parts of the flower are most frequently arranged in fives, or multiples of fives; for instance, a common arrangement is as follows, - five sepals, succeeded by five petals, ten stamens in two sets of five, and five or fewer carpels; an arrangement in fours is less frequent, while the arrangement in threes, so common in monocotyledons, is rare in dicotyledons.
In some orders the parts are numerous, chiefly in the case of the stamens and the carpels, as in the buttercup and other members of the order Ranunculaceae.
There is a very wide range in the general structure and arrangement of the parts of the flower, associated with the means for ensuring the transference of pollen; in the simplest cases the flower consists only of a few stamens or carpels, with no enveloping sepals or petals, as in the willow, while in, the more elaborate type each series is represented, the whole forming a complicated structure closely correlated with the size, form and habits of the pollinating agent (see Flower).
These stamens encircle a style which is the upward continuation of the ovary, and which shows at its free end traces of the three originally separate but now blended carpels of which the ovary consists.
In both cases the socalled fruit is composed of the receptacle or upper end of the flower-stalk (the so-called calyx tube) greatly dilated, and enclosing within its cellular flesh the five cartilaginous carpels which constitute the "core" and are really the true fruit.
The ovary, of two carpels, is seated on a ring-like disk FIG.
And Sambucus, more rarely two-lipped as in Lonicera; the sepals and petals are usually five in number and placed above the ovary, the five stamens are attached to the corolla-tube, there are three to five carpels, and the fruit is a berry as in honeysuckle or snowberry (Symphoricarpus), or a stone fruit, with several, usually three, stones, as in Sambucus.
The pistil, which is above the rest of the members of the flower, consists of two carpels joined at their edges to form the ovary, which becomes two-celled by subsequent ingrowth of a septum from these united edges; a row of ovules springs from each edge.
After the carpels have fallen away.
5), the perianth which is generally petaloid occupying the two outer whorls, followed by two whorls of stamens, with a superior ovary of three carpels in the centre of the flower; the ovary is generally three-chambered and contains an indefinite number of anatropous ovules on axile placentas (see fig.
The parts of the flower are in fives in calyx, corolla and stamens, followed by two carpels which unite to form a superior ovary.
The female flowers are equally simple, consisting of a bract, from whose axil arises usually a very short stalk, surmounted by two carpels adherent one to the other for their whole length, except that the upper ends of the styles are separated into two stigmas.
When ripe the two carpels separate in the form of two valves and liberate a large number of seeds, each provided at the base with a tuft of silky hairs, and containing a straight embryo without any investing albumen.
The three carpels forming the pepo are separated by partitions (cl).
The sepals and petals are free or more or less united, the stamens as many or twice as many as the petals; the carpels, usually free, are equal to the petals in number, and form in the fruit follicles with two or more seeds.
The structure of the flower represents the simple type of monocotyledons, consisting of two whorls of petals, of three free parts each, six free stamens, and a consolidated pistil of three carpels, ripening into a three-valved capsule containing many winged seeds.
Xenogamy is of course the only possible method in diclinous plants; it is also the usual method in monoclinous plants, owing to the fact that stamens and carpels often mature at different times (dichogamy), the plants being proterandrous or proterogynous.
There are five, or sometimes fewer, carpels, which unite to form an ovary with as many chambers, in each of which are one or two, rarely more, pendulous anatropous ovules, attached to the central column in such a way that the micropyle points outwards and the raphe is turned towards the placenta.
In the ripe fruit the carpels separate into five one-seeded portions (cocci), which break away from the central column, either rolling elastically outwards and upwards or becoming spirally twisted.
The numerous stamens surround the ovary, which is composed of 4 to 16 carpels and is surmounted by a flat or convex rayed disk bearing the stigmas.
The carpel, or aggregate of carpels forming the pistil or gynaeceum, comprises an ovary containing one or more ovules and a receptive surface or stigma; the stigma is sometimes carried up on a style.
In Gymnosperms we have seeds, and the carpels may become modified and close around these, as in Pinus, during the process of ripening to form an imitation of a box-like fruit which subsequently opening allows the seeds to escape; but there is never in them the closed ovary investing from the outset the ovules, and ultimately forming the ground-work of the fruit.
Series 6, Apocarpeae, is characterized by 5 carpels, and in the last series Glumaceae, great simplification in the flower is associated with a grass-like habit.
In diameter, with five spreading white petals alternating with five persistent green sepals, a large number of stamens with pinkish-brown anthers, and one to three carpels sunk in the cup-shaped floral axis.
The irregular flowers have five sepals united at the base, the dorsal one produced into a spurred development of the axis; of the five petals the two upper are slightly different and stand rather apart from the lower three; the eight stamens are unequal and the pistil consists of three carpels which form a fleshy fruit separating into three one-seeded portions.
Double nuts are the result of the equal development of the two carpels of the original flower, of which ordinarily one becomes abortive; fusion of two or more nuts is not uncommon.
In a few cases two whorls of stamens are present, with three members in each, but generally only three are present; the pistil consists of three or two carpels, united to form an ovary bearing a corresponding number of styles and containing one ovule.
R 5), it will be seen to differ in the absence of the outer row and the posterior member of the inner row of the perianth-leaves, of the whole inner row of stamens, and of the two lateral carpels, FIG.
Dioecious; flowers in the form of cones, except the female flowers of Cycas, which consist of a rosette of leaf-like carpels at the apex of the stem.
In the genus Cycas the female flower is peculiar among cycads in consisting of a terminal crown of separate leaf-like carpels several inches in length; the apical portion of each carpellary leaf may be broadly triangular in form, and deeply dissected on the margins into narrow woolly appendages like rudimentary pinnae.
From the lower part of a carpel are produced several laterally placed ovules, which become bright red or orange on ripening; the bright fleshy seeds, which in some species are as large as a goose's egg, and the tawny spreading carpels produce a pleasing combination of colour in the midst of the long dark-green fronds, which curve gracefully upwards and outwards from the summit of the columnar stem.
I) on the branched or unbranched stem; (b) the growth of the main stem through the female flower; (c) the presence of a prominent single vein in the linear pinnae; (d) the structure of the female flower, which is peculiar in not having the form of a cone, but consists of numerous independent carpels, each of which bears two or more lateral ovules.
- Like Zamia, except that the ends of the stamens are flat, while the apices of the carpels are peltate.
- Characterized by the woolly scaleleaves and carpels; the latter terminate in a thick laminar expansion of triangular form, bearing two placental cushions, on which the ovules are situated.
In Cycas revoluta and C. circinalis each leaf-like carpel may produce several laterally attached ovules, but in C. Normanbyana the carpel is shorter and the ovules are reduced to two; this latter type brings us nearer to the carpels of Dioon, in which the flower has the form of a cone, and the distal end of the carpels is longer and more leaf-like than in the other genera of the Zamieae, which are characterized by shorter carpels with thick peltate heads bearing two ovules on the morphologically lower surface.
These projections and ridges may be homologous with the seminiferous scale of the pines, firs, cedars, &c. The simplest interpretation of the cone of the Abietineae is that which regards it as a flower consisting of an axis bearing several open carpels, which in the adult cone may be very small or large and prominent, the scale bearing the ovules being regarded as a placental outgrowth from the flat and open carpel.
Robert Brown was the first to give a clear description of the morphology of the Abietineous cone in which carpels bear naked ovules; he recognized gymnospermy as an important distinguishing feature in conifers as well as in cycads.