All the incidents subsequent to the appeal of Athens to Sparta are expressly referred by Herodotus to the interval between the sending of the heralds in 491 B.C. and the invasion of Datis and Artaphernes in 490 B.C. (cf.
ARTAPHERNES, more correctly Artaphrenes, brother of Darius Hystaspis, and satrap of Sardis.
In 494, when the defeated Ionians were obliged to sue for terms, he was one of the ambassadors to the Persian satrap Artaphernes, whom he persuaded to restore the constitution of the Ionic cities (Diod.
It was in consequence of the aid which the people of Miletus lent to the Eretrians on this occasion that Eretria sent five ships to aid the Ionians in their revolt against the Persians (see IoNiA); and owing to this, that city was the first place in Greece proper to be attacked by Datis and Artaphernes in 490 B.C. It was utterly ruined on that occasion, and its inhabitants were transported to Persia.