Formation of archenteron and blastopore may, however, be deferred till a later stage (actinula or after).
Some form their diblastula by emboly, others by epiboly; and in the later history of the further development of the enclosed cells (archenteron) very marked variations occur in closely-allied forms, due to the influence of a greater or less abundance of food-material mixed with the protoplasm of the egg.
In the direct development Bateson showed that the three divisions of the coelom arise as pouches constricted off from the archenteron or primitive gut, thus resembling the development of the mesoblastic somites of Amphioxus.
Segmentation is complete, a gastrula is formed, the blastopore closes, the archenteron gives off two coelomic sacs which, as far as is known, are unaffected by the super ficial segmentation of the body that divides the larva into three segments.
Of these latter, two grades were further distinguished by Lankester - those which remain possessed of a single archenteric cavity and of two primary cell-layers (the Coelentera or Diploblastica), and those which by nipping off the archenteron give rise to two cavities, the coelom or body-cavity and the metenteron or gut (Coelomata or Triploblastica).
In this stage the body is composed of two layers, ectoderm (d) externally, and endoderm (c) internally, surrounding a central cavity, the archenteron (b), which communicates with the exterior by a pore (a), the blastopore.
The archenteron becomes the gastrovascular system or coelenteron.
In common with all other Coelomata, the Mollusca have the mouth and first part of the alimentary canal which leads into the met-enteron formed by a special invagination of the outer layer of the primitive body-wall, not to be confounded with that which often, but not always, accompanies the antecedent formation of the archenteron; this invagination is termed the stomodaeum.
The archenteron gives off two lateral pounchs and thus becomes trilobed.
The embryo now consists of two layers of cells, epiblast and hypoblast, surrounding a cavity, the archenteron, which opens to the exterior by the orifice of invagination or blastopore.
An important fact to note is that the blastopore is included in this overgrowth of epiblast, so that the neural tube remains for some time in open communication with the archenteron by means of a posterior neurenteric canal.
In front of the latter there remains a portion of the archenteron, which becomes constricted off as the head cavity.
It is reasonable to suppose that the Coelomata - animals in which the body-cavity is divided into a gut passing from mouth to anus and a hollow (coelom) surrounding it - were derived from the simpler Coelentera, in which the primitive body-cavity (archenteron) is not so divided, and has only one aperture serving as both mouth and anus.
We may, with Sedgwick, suppose the coelom to have originated by the enlargement and separation of pouches that pressed outwards from the archenteron into the thickened body-wall (such structures as the genital pouches of some Coelentera, not yet shut off from the rest of the cavity), and they would probably have been four in number and radially disposed about the central cavity.